Thomisidae: Xysticus C. L. Koch, 1835

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1
Males

  2  

- Females

  22  

2
(1)
Median bulbal apophysis T-shaped; ventral tibial apophysis broadly truncated

  3  

- Different

  5  

3
(2)

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Tip of embolus broad, flanked by 2 wing-like structures which are transparent distally

Xysticus macedonicus Šilhavý, 1944

female

- Embolic tip different

  4  

4
(3)

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cp

Basal part of T-shaped bulbal apophysis longer than wide

Xysticus cristatus (Clerck, 1757)

female

-

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Basal part of T-shaped bulbal apophysis ± as long as wide

Xysticus audax (Schrank, 1803)

female

5
(2)
At least one bulbal apophysis robust, transverse to longitudinal bulbal axis, approx. 1/2 of bulbus width

  6  

- Different

  11  

6
(5)

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Median bulbal apophysis with 2 tips, arranged rectangularly

Xysticus luctator L. Koch, 1870

female

- Different

  7  

7
(6)

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Both bulbal apophyses ± parallel and equal in length, pointed

Xysticus bifasciatus C. L. Koch, 1837

female

- Bulbal apophysis different

  8  

8
(7)
Bulbus with two transverse apophysis

  9  

- Bulbus with one transverse apophysis

  10  

9
(8)

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Bulbal apophysis slightly bent toward each other, pliers-like

Xysticus kochi Thorell, 1872

female

-

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One bulbal apophysis hammer-shaped

Xysticus obscurus Collett, 1877

female

10
(8)

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Bulbal apophysis as fig.

Xysticus erraticus (Blackwall, 1834)

female

-

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Both bulbal apophyses pointing in opposite direction

Xysticus viduus Kulczyński, 1898

female

11
(5)
Bulbus medially with 2 short, robust apophyses, directing pliers-like toward each other

  12  

- Bulbus without such apophysis

  13  

12
(11)

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cp

Both pliers-like apophyses bent

Xysticus ulmi (Hahn, 1831)

female

-

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Only one of these apophyses bent

Xysticus lanio C. L. Koch, 1835

female

13
(11)

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cp

Tip of embolus strikingly broad and with transverse folds

Spiracme striatipes (L. Koch, 1870)

female

- Different

  14  

14
(13)
Lateral tibial apophysis with pointed ending

  15  

- Lateral tibial apophysis rounded

  19  

15
(14)

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Tip of lateral tibial apophysis strikingly narrow and elongated

Xysticus lineatus (Westring, 1851)

female

- Different

  16  

16
(15)
Lateral tibial apophysis ± triangular

  17  

- Lateral tibial apophysis with ± parallel sides

  18  

17
(16)

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Ventral tibial apophysis with 2 distinctly separated processes

Xysticus kempeleni Thorell, 1872

female

-

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The two processes of ventral tibial apophysis basally merged

Xysticus luctuosus (Blackwall, 1836)

female

18
(16)

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Tip of lateral tibial apophysis directing ventrally; ventral tibial apophysis small

Xysticus acerbus Thorell, 1872

female

-

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Tip of lateral tibial apophysis directing dorsally; ventral tibial apophysis large and with several processes

Bassaniodes robustus (Hahn, 1832)

female

19
(14)
Cymbial process strikingly broad basally or bent

  20  

- Cymbial process small

  21  

20
(19)

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Cymbial process basally broad; tip of embolus thin, distally twisted

Psammitis bonneti (Denis, 1938)

female

-

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Cymbial process S-like bent; tip of embolus broad

Psammitis secedens (L. Koch, 1876)

female

21
(19)

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Lateral tibial apophysis basally wider than distally; tip of embolus bent

Psammitis sabulosus (Hahn, 1832)

female

-

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Lateral tibial apophysis with approx. parallel sides; tip of embolus straight

Psammitis ninnii (Thorell, 1872)

female

22
(1)
Epigyne with two grooves which are distinctly separated

  33  

- Epigyne with one groove

  23  

23
(22)
Distance between epigastric furrow and posterior margin of epigynal groove smaller or equal one diameter of epigynal groove

  24  

- Distance between epigastric furrow and posterior margin of epigynal groove 1.5-2 diameters of epigynal grooves

  31  

24
(23)
Epigynal groove medially indented on anterior margin

  25  

- Different

  26  

25
(24)

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Epigyne as fig.

Psammitis ninnii (Thorell, 1872)

male

-

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Epigyne as fig.

Xysticus viduus Kulczyński, 1898

male

26
(24)

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Epigynal groove posteriorly distinctly narrower than anteriorly

Xysticus luctuosus (Blackwall, 1836)

male

- Different

  27  

27
(26)
Epigynal groove ± quadrangular or triangular

  28  

- Epigynal groove roundish

  30  

28
(27)

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Epigynal groove ± triangular, particularly laterally bordered by robust chitinous wall

Psammitis bonneti (Denis, 1938)

male

- Epigynal groove ± quadrangular

  29  

29
(28)

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Epigynal groove longish, ± quadrangular, anteriorly bordered by broad chitinous wall

Xysticus lineatus (Westring, 1851)

male

-

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Epigynal groove quadratic to roundish

Xysticus obscurus Collett, 1877

male

30
(27)

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All sides of epigynal groove distinctly bordered

Xysticus acerbus Thorell, 1872

male

-

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Epigynal groove indistinctly bordered anteriorly

Psammitis sabulosus (Hahn, 1832)

male

31
(23)

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Basic shape of epigyne roundish

Xysticus bifasciatus C. L. Koch, 1837

male

- Basic shape of epigyne rhombic/trapezoid

  32  

32
(31)

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Epigynal groove laterally indented; vulva as fig.

Xysticus erraticus (Blackwall, 1834)

male

-

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Epigynal groove laterally not indented; vulva as fig.

Bassaniodes robustus (Hahn, 1832)

male

33
(22)

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Epigyne ± quadrangular with rounded corners; margins of bar between the two epigynal grooves anteriorly diverging

Xysticus kempeleni Thorell, 1872

male

- Different

  34  

34
(33)

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Bar between the two epigynal grooves distinctly wider than width of one epigynal groove

Xysticus luctator L. Koch, 1870

male

- Different

  35  

35
(34)
Epigyne anteriorly with broad, semicircular chitinous wall

  36  

- Different

  37  

36
(35)

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Width of bar between the two epigynal grooves approx. width of epigynal groove

Spiracme striatipes (L. Koch, 1870)

male

-

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Width of bar between the two epigynal grooves approx. 1/2 width of epigynal groove

Xysticus kochi Thorell, 1872

male

37
(35)

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Inner margins of epigynal groove strongly sclerotised medially

Xysticus audax (Schrank, 1803)

male

- Different

  38  

38
(37)
Epigynal region all in all more strongly sclerotised and ± roundish

  39  

-

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Epigynal region only at outer sides of epigynal grooves more strongly sclerotised, so that it consists of two separated parts

Psammitis secedens (L. Koch, 1876)

male

39
(38)

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Bar between the epigynal grooves ± as wide as one epigynal groove

Xysticus lanio C. L. Koch, 1835

male

- Bar between epigynal grooves distinctly narrower than one epigynal groove

  40  

40
(39)

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Epigyne and vulva as fig.

Xysticus ulmi (Hahn, 1831)

male

- Different

  41  

41
(40)

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Posterior margin of epigynal grooves distinctly sclerotised; Vulva as fig.

Xysticus cristatus (Clerck, 1757)

male

-

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Posterior margin of epigynal grooves less sclerotised; vulva as fig.

Xysticus macedonicus Šilhavý, 1944

male

Further taxa

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1. Xysticus abditus Logunov, 2006

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2. Xysticus alpicola Kulczyński, 1882

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3. Xysticus alpinus Kulczyński, 1887

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4. Xysticus apricus L. Koch, 1876

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5. Xysticus atevs Ovtsharenko, 1979

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6. Xysticus austrosibiricus Logunov & Marusik, 1998

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7. Xysticus bacurianensis Mcheidze, 1971

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8. Xysticus bicolor L. Koch, 1867

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9. Xysticus brevidentatus Wunderlich, 1995

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10. Xysticus britcheri Gertsch, 1934

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11. Xysticus canadensis Gertsch, 1934

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12. Xysticus caspicus Utochkin, 1968

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13. Xysticus caucasius L. Koch, 1878

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14. Xysticus charitonowi Mcheidze, 1971

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15. Xysticus chippewa Gertsch, 1953

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16. Xysticus cor Canestrini, 1873

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17. Xysticus corsicus Simon, 1875

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18. Xysticus desidiosus Simon, 1875

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19. Xysticus diversus (Blackwall, 1870)

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20. Xysticus edax (O. Pickard-Cambridge, 1872)

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21. Xysticus emertoni Keyserling, 1880

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22. Xysticus ferrugineus Menge, 1876

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23. Xysticus gallicus Simon, 1875

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24. Xysticus gortanii Caporiacco, 1922

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25. Xysticus grallator Simon, 1932

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26. Xysticus ibex Simon, 1875

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27. Xysticus idolothytus Logunov, 1995

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28. Xysticus kalandadzei Mcheidze & Utochkin, 1971

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29. Xysticus kaznakovi Utochkin, 1968

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30. Xysticus kulczynskii Werjbitzky, 1902

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31. Xysticus laetus Thorell, 1875

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32. Xysticus lalandei (Audouin, 1826)

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33. Xysticus lendli Kulczyński, 1897

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34. Xysticus marmoratus Thorell, 1875

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35. Xysticus marusiki Ono & Martens, 2005

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36. Xysticus metinaktasi Demir, Seyyar & Türker, 2017

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37. Xysticus mongolicus Schenkel, 1963

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38. Xysticus nataliae Utochkin, 1968

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39. Xysticus nigrotrivittatus (Simon, 1870)

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40. Xysticus nubilus Simon, 1875

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41. Xysticus ovatus Simon, 1876

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42. Xysticus parallelus Simon, 1873

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43. Xysticus pseudolanio Wunderlich, 1995

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44. Xysticus pseudoluctuosus Marusik & Logunov, 1995

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45. Xysticus semicarinatus Simon, 1932

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46. Xysticus siciliensis Wunderlich, 1995

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47. Xysticus sjostedti Schenkel, 1936

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48. Xysticus slovacus Svaton, Pekar & Pridavka, 2000

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49. Xysticus soderbomi Schenkel, 1936

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50. Xysticus spasskyi Utochkin, 1968

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51. Xysticus tenuiapicalis Demir, 2012

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52. Xysticus thessalicoides Wunderlich, 1995

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53. Xysticus thessalicus Simon, 1916

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54. Xysticus tortuosus Simon, 1932

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55. Xysticus ukrainicus Utochkin, 1968

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56. Xysticus xerodermus Strand, 1913

world57. Xysticus acerbus obscurior Kulczyński, 1895
world58. Xysticus arenarius Thorell, 1875
world59. Xysticus arenicola Simon, 1875
world60. Xysticus audax massanicus Simon, 1932
world61. Xysticus barbatus Caporiacco, 1936
world62. Xysticus clercki (Audouin, 1826)
world63. Xysticus doriai (Dalmas, 1922)
world64. Xysticus ibex dalmasi Simon, 1932
world65. Xysticus kempeleni nigriceps Simon, 1932
world66. Xysticus peccans O. Pickard-Cambridge, 1876
world67. Xysticus promiscuus O. Pickard-Cambridge, 1876
world68. Xysticus quadrispinus Caporiacco, 1933