Gnaphosidae: Haplodrassus Chamberlin, 1922

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1
 Males

  2  
- Females

  12  
2
(1)

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cp

 Tibial apophysis narrow; body length smaller or equal 4.0 mm; prosoma length smaller or equal 2.0 mm

Haplodrassus minor (O. Pickard-Cambridge, 1879)

female
speciesimage

male Pedipalp, ventral view
(Locket et al. 1974)

 speciesimage

male Pedipalp, retrolateral view
(Roberts 1995)

 speciesimage

male Palpal tibia
(Locket et al. 1974)
speciesimage

male Tibial apophysis
(Miller & Buchar 1977)
- Tibial apophysis different and/or body length larger

  3  
3
(2)		 Tibial apophysis distally broadened

  4  
- Tibial apophysis distally not broadened

  8  
4
(3)

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cp

 Distal bulbal apophysis with strikingly serrated margin

Haplodrassus kulczynskii Lohmander, 1942

female
speciesimage

male Pedipalp, ventral view
(Miller 1971)

 speciesimage

male Pedipalp, retrolateral view
(Roberts 1998)

 speciesimage

male Tibial apophysis
(Miller 1971)
- Distal bulbal apophysis different

  5  
5
(4)		 Distal bulbal apophysis long and slender

  6  
- Distal bulbal apophysis broader and

  9  
6
(5)

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cp

 Pedipalp as fig.; tibial apophysis distally hardly broadened

Haplodrassus bohemicus Miller & Buchar, 1977

female
speciesimage

male Pedipalp, ventral view
(Miller & Buchar 1977)

 speciesimage

male Embolus
(Miller & Buchar 1977)

 speciesimage

male Palpal tibia, dorsal view
(Miller & Buchar 1977)
- Different

  7  
7
(6)

world

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cp

 Distal bulbal apophysis distinctly keeled

Haplodrassus signifer (C. L. Koch, 1839)

female
speciesimage

male Pedipalp, ventral view
(Brændegaard 1966)

 speciesimage

male Pedipalp, ventral view
(Noordam 1992)

 speciesimage

male Pedipalp, retrolateral view
(Noordam 1992)
speciesimage

male Pedipalp, retrolateral view
(Roberts 1995)

 speciesimage

male Pedipalp, retrolateral view
(Locket & Millidge 1951)

 speciesimage

male Tip of pedipalp
(Locket & Millidge 1951)
speciesimage

male Tibial apophysis
(Brændegaard 1966)

 speciesimage

male Tibial apophysis
(Noordam 1992)

 speciesimage

male Tibial apophysis
(Locket & Millidge 1951)
-

world

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cp

 Distal bulbal apophysis without keel

Haplodrassus aenus Thaler, 1984

female
speciesimage

male Pedipalp, ventral view
(Thaler 1984a)

 speciesimage

male Tibial apophysis
(Thaler 1984a)
8
(3)

world

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cp

 Plate-like bulge of distal bulbal apophysis situated distally, overtops tip of apophysis; tibial apophysis distinctly broadened distally

Haplodrassus soerenseni (Strand, 1900)

female
speciesimage

male Pedipalp, ventral view
(Miller 1971)

 speciesimage

male Pedipalp, retrolateral view
(Roberts 1995)

 speciesimage

male Pedipalp, retrolateral view
(Locket & Millidge 1953)
speciesimage

male Tip of pedipalp, ventral view
(Locket et al. 1974)

 speciesimage

male Embolic division, ventral view
(Locket & Millidge 1953)

 speciesimage

male Tibial apophysis
(Miller & Buchar 1977)
-

world

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cp

 Plate-like bulge of distal bulbal apophysis situated basally; tibia apophysis only little broadened

Haplodrassus umbratilis (L. Koch, 1866)

female
speciesimage

male Pedipalp, ventral view
(Miller 1971)

 speciesimage

male Pedipalp, ventral view
(Locket et al. 1974)

 speciesimage

male Pedipalp, retrolateral view
(Roberts 1995)
speciesimage

male Palpal tibia
(Locket et al. 1974)

 speciesimage

male Tibial apophysis
(Miller & Buchar 1977)
9
(5)		 Distal bulbal apophysis distinctly broadened distally

  10  
-

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cp

 Distal bulbal apophysis not broadened distally and with 2 tooth-like structures

Haplodrassus dalmatensis (L. Koch, 1866)

female
speciesimage

male Pedipalp, ventral view
(Noordam 1992)

 speciesimage

male Pedipalp, ventral view
(Locket et al. 1974)

 speciesimage

male Pedipalp, retrolateral view
(Noordam 1992)
speciesimage

male Pedipalp, retrolateral view
(Roberts 1995)

 speciesimage

male Pedipalp, retrolateral view
(Locket & Millidge 1951)

 speciesimage

male Tibial apophysis
(Noordam 1992)
speciesimage

male Tibial apophysis
(Locket & Millidge 1951)
10
(9)

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cp

 Distal bulbal apophysis ± straight, basally with a tooth; tibial apophysis as fig.

Haplodrassus cognatus (Westring, 1861)

female
speciesimage

male Palpal tibia, dorsal view
(Miller & Buchar 1977)
- Distal bulbal apophysis ± rectangularly bent, without tooth

  11  
11
(10)

world

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cp

 Tibial apophysis large, reaching 1/2 of bulbal length, distinctly keeled

Haplodrassus silvestris (Blackwall, 1833)

female
speciesimage

male Pedipalp, ventral view
(Brændegaard 1966)

 speciesimage

male Pedipalp, ventral view
(Noordam 1992)

 speciesimage

male Pedipalp, retrolateral view
(Noordam 1992)
speciesimage

male Pedipalp, retrolateral view
(Roberts 1995)

 speciesimage

male Pedipalp
(Locket & Millidge 1951)

 speciesimage

male Tip of pedipalp
(Locket & Millidge 1951)
speciesimage

male Tibial apophysis
(Noordam 1992)

 speciesimage

male Tibial apophysis
(Locket & Millidge 1951)
-

world

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cp

 Tibial apophysis small, reaching about 1/3 of bulbal length, without distinct keel

Haplodrassus moderatus (Kulczyński, 1897)

female
speciesimage

male Pedipalp, ventral view
(Miller 1971)

 speciesimage

male Pedipalp, retrolateral view
(Roberts 1998)
12
(1)		 Epigyne with several transverse folds

  13  
- Different

  14  
13
(12)

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cp

 Bar of anterior epigynal margin narrow; depressed area between the sclerotised pockets narrow

Haplodrassus silvestris (Blackwall, 1833)

male
speciesimage

male Tip of pedipalp
(Locket & Millidge 1951)

 speciesimage

female Epigyne
(Brændegaard 1966)

 speciesimage

female Epigyne
(Noordam 1992)
speciesimage

female Epigyne
(Roberts 1995)

 speciesimage

female Epigyne
(Locket & Millidge 1951)

 speciesimage

female Vulva
(Miller & Buchar 1977)
speciesimage

female Vulva, ventral view
(Tullgren 1946)
-

world

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cp

 Bar of anterior epigynal margin narrow as broad as epigyne; depressed area between sclerotised pockets broad and with strong indentations

Haplodrassus cognatus (Westring, 1861)

male
speciesimage

female Epigyne
(Brændegaard 1966)

 speciesimage

female Epigyne
(Miller 1971)

 speciesimage

female Vulva
(Miller & Buchar 1977)
speciesimage

female Vulva, ventral view
(Tullgren 1946)
14
(12)		 Sclerotised epigynal pockets with shallow protrusion

  15  
- Sclerotised epigynal pockets without protrusion

  17  
15
(14)

world

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cp

 Protrusion very elongated

Haplodrassus kulczynskii Lohmander, 1942

male
speciesimage

female Epigyne
(Miller 1971)

 speciesimage

female Epigyne
(Roberts 1998)

 speciesimage

female Vulva
(Miller & Buchar 1977)
- Different

  16  
16
(15)

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cp

 Protrusion slantingly forward-directed

Haplodrassus umbratilis (L. Koch, 1866)

male
speciesimage

female Epigyne
(Brændegaard 1966)

 speciesimage

female Epigyne
(Roberts 1995)

 speciesimage

female Epigyne
(Locket et al. 1974)
speciesimage

female Vulva
(Miller & Buchar 1977)

 speciesimage

female Vulva, ventral view
(Tullgren 1946)
-

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cp

 Protrusions directed more towards median region

Haplodrassus soerenseni (Strand, 1900)

male
17
(14)		 Margins of sclerotised, black epigynal pockets parallel or slightly roundish curved

  18  
- Margins of sclerotised, black epigynal pockets strongly converging or diverging posteriorly

  21  
18
(17)		 Margins on the inner side roundish curved

  19  
- Margins on the inner side straight

  20  
19
(18)

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cp

 Epigyne and vulva as figs.

Haplodrassus signifer (C. L. Koch, 1839)

male
speciesimage

female Epigyne
(Brændegaard 1966)

 speciesimage

female Epigyne
(Noordam 1992)

 speciesimage

female Epigyne
(Roberts 1995)
speciesimage

female Epigyne
(Locket & Millidge 1951)

 speciesimage

female Vulva
(Platnick & Shadab 1975)

 speciesimage

female Vulva, ventral view
(Tullgren 1946)
-

world

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cp

 Epigyne and vulva as figs.

Haplodrassus bohemicus Miller & Buchar, 1977

male
speciesimage

female Epigyne
(Miller & Buchar 1977)

 speciesimage

female Vulva
(Miller & Buchar 1977)
20
(18)

world

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cp

 Epigyne and vulva as figs.

Haplodrassus aenus Thaler, 1984

male
speciesimage

female Epigyne/vulva, ventral view
(Thaler 1984a)

 speciesimage

female Epigyne/vulva, ventral view
(Thaler 1984a)
-

world

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cp

 Epigyne and vulva as figs.

Haplodrassus minor (O. Pickard-Cambridge, 1879)

male
speciesimage

female Epigyne
(Roberts 1995)

 speciesimage

female Epigyne
(Locket & Millidge 1951)
21
(17)

world

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cp

 Margins diverging towards posterior

Haplodrassus dalmatensis (L. Koch, 1866)

male
speciesimage

female Epigyne
(Miller 1971)

 speciesimage

female Epigyne
(Brændegaard 1966)

 speciesimage

female Epigyne
(Miller 1971)
speciesimage

female Epigyne
(Noordam 1992)

 speciesimage

female Epigyne
(Roberts 1995)

 speciesimage

female Epigyne
(Locket & Millidge 1951)
speciesimage

female Vulva
(Miller & Buchar 1977)
-

world

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cp

 Margins converging towards posterior

Haplodrassus moderatus (Kulczyński, 1897)

male
speciesimage

female Epigyne
(Roberts 1998)

 speciesimage

female Vulva, ventral view
(Tullgren 1946)

Further taxa

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1. Haplodrassus alexeevi Ponomarev & Shmatko, 2017
speciesimage

male Pedipalp, ventral view
(Ponomarev & Shmatko 2017)

 speciesimage

male Bulbus, ventral view
(Ponomarev & Shmatko 2017)

 speciesimage

male Bulbus, ventro-lateral view
(Ponomarev & Shmatko 2017)
speciesimage

male Tibial apophysis, dorsal view
(Ponomarev & Shmatko 2017)

 speciesimage

female Epigyne
(Ponomarev & Shmatko 2017)

 speciesimage

female Vulva
(Ponomarev & Shmatko 2017)

world

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cp

2. Haplodrassus caspius Ponomarev & Belosludtsev, 2008
speciesimage

male Pedipalp, retrolateral view
(Kovblyuk et al. 2013d)

 speciesimage

male Pedipalp, ventral view
(Kovblyuk et al. 2013d)

 speciesimage

male Pedipalp, prolateral view
(Kovblyuk et al. 2013d)
speciesimage

male Pedipalp, retrolateral view
(Chatzaki 2021)

 speciesimage

male Pedipalp, ventral view
(Chatzaki 2021)

 speciesimage

male Bulbus, dorsal view
(Kovblyuk et al. 2013d)
speciesimage

male Bulbus, apical view
(Kovblyuk et al. 2013d)

 speciesimage

male Tip of bulbus, dorsal view
(Chatzaki 2021)

 speciesimage

female Epigyne, ventral view
(Ponomarev et al. 2008)
speciesimage

female Epigyne, ventral view
(Kovblyuk et al. 2013d)

 speciesimage

female Vulva, dorsal view
(Ponomarev et al. 2008)

 speciesimage

female Vulva, dorsal view
(Kovblyuk et al. 2013d)
speciesimage

female Eyes, frontal view
(Kovblyuk et al. 2013d)

 speciesimage

female Chelicera, posterior view
(Kovblyuk et al. 2013d)

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cp

3. Haplodrassus caucasius Ponomarev & Dvadnenko, 2013
speciesimage

male Pedipalp, ventral view
(Ponomarev & Dvadnenko 2013)

 speciesimage

male Pedipalp, retrolateral view
(Ponomarev & Dvadnenko 2013)

 speciesimage

female Epigyne, ventral view
(Ponomarev & Dvadnenko 2013)

world

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cp

4. Haplodrassus concertor (Simon, 1878)
speciesimage

male Pedipalp, prolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)
speciesimage

male Apical part of bulbus
(Simon 1914)

 speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Simon 1914)
speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)

 speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

world

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cp

5. Haplodrassus crassipes (Lucas, 1846)
speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Embolic apophyis, ventral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)
speciesimage

male Palpal tibia, retrolateral view
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Bosmans et al. 2018)
speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)

 speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

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6. Haplodrassus creticus (Roewer, 1928)
speciesimage

male Pedipalp, ventral view
(Chatzaki et al. 2002a)

 speciesimage

male Pedipalp, retrolateral view
(Chatzaki et al. 2002a)

 speciesimage

male Pedipalp, prolateral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Bulbus
(Chatzaki et al. 2002a)
speciesimage

male Palpal tibia, dorsal view
(Chatzaki et al. 2002a)

 speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Chatzaki et al. 2002a)
speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Vulva
(Chatzaki et al. 2002a)

 speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

world

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7. Haplodrassus dentifer Bosmans & Abrous, 2018
speciesimage

male Pedipalp, prolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Embolic division, ventral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)
speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Bosmans et al. 2018)
speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

 speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)

world

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8. Haplodrassus ibericus Melic, Silva & Barrientos, 2016
speciesimage

male Pedipalp, ventro-apical view
(Melic et al. 2016)

 speciesimage

male Pedipalp, prolateral view
(Melic et al. 2016)

 speciesimage

male Pedipalp, retrolateral view
(Melic et al. 2016)
speciesimage

male Pedipalp, ventral view
(Melic et al. 2016)

 speciesimage

male Tibial apophysis, dorsal view
(Melic et al. 2016)

 speciesimage

female Epigyne
(Melic et al. 2016)
speciesimage

female Vulva
(Melic et al. 2016)

world

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9. Haplodrassus invalidus (O. Pickard-Cambridge, 1872)
speciesimage

male Pedipalp, ventral view
(Levy 2004)

 speciesimage

male Pedipalp, ventral view
(Kovblyuk et al. 2012b)

 speciesimage

male Pedipalp, prolateral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Bulbus, apical view
(Kovblyuk et al. 2012b)
speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)

 speciesimage

male Tibial apophysis
(Levy 2004)

 speciesimage

male Tibial apophysis, retrolateral view
(Kovblyuk et al. 2012b)
speciesimage

female Epigyne
(Levy 2004)

 speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Chatzaki & Van Keer 2019)
speciesimage

female Vulva, dorsal view
(Levy 2004)

 speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

 speciesimage

female Vulva
(Chatzaki & Van Keer 2019)

world

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10. Haplodrassus ivlievi Ponomarev, 2015
speciesimage

female Epigyne, ventral view
(Ponomarev 2015)

 speciesimage

female Vulva, dorsal view
(Ponomarev 2015)

world

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11. Haplodrassus longivulva Bosmans & Hervé, 2018
speciesimage

male Pedipalp, prolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)
speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)
speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

world

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12. Haplodrassus lyndae Abrous & Bosmans, 2018
speciesimage

male Pedipalp, prolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Breitling 2020)

 speciesimage

male Embolus, dorsal view
(Bosmans et al. 2018)
speciesimage

male Embolic division, ventral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, retrolateral view
(Breitling 2020)

 speciesimage

male Tibial apophysis, dorso-lateral view
(Bosmans et al. 2018)
speciesimage

male Tibial apophysis, dorso-lateral view
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Bosmans et al. 2018)
speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)

 speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

 speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)

world

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13. Haplodrassus macellinus (Thorell, 1871)
speciesimage

male Pedipalp, ventral view
(Di Franco 1996)

 speciesimage

male Pedipalp
(Di Franco 1996)

 speciesimage

male Pedipalp, ventral view
(Simon 1878)
speciesimage

male Pedipalp, retrolateral view
(Lecigne unpubl.)

 speciesimage

male Pedipalp, ventral view
(Lecigne unpubl.)

 speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, retrolateral view
(Lecigne 2018b)

 speciesimage

male Pedipalp, ventral view
(Lecigne 2018b)
speciesimage

male Pedipalp, ventro-prolateral view
(Lecigne 2018b)

 speciesimage

male Palpal tibia
(O. Pickard-Cambridge 1874)

 speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)
speciesimage

male Palpal tibia, retrolateral view
(Lecigne 2018b)

 speciesimage

female Epigyne
(Di Franco 1996)

 speciesimage

female Epigyne, ventral view
(Simon 1878)
speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Vulva
(Di Franco 1996)

 speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)
speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

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14. Haplodrassus mediterraneus Levy, 2004
speciesimage

male Pedipalp, ventral view
(Levy 2004)

 speciesimage

male Pedipalp, prolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, retrolateral view
(Levy 2004)

 speciesimage

female Epigyne, ventral view
(Levy 2004)
speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Vulva, dorsal view
(Levy 2004)

 speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)
speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

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15. Haplodrassus morosus (O. Pickard-Cambridge, 1872)
speciesimage

male Pedipalp, ventral view
(Levy 2004)

 speciesimage

male Tibial apophysis
(Levy 2004)

 speciesimage

female Epigyne
(Levy 2004)
speciesimage

female Vulva
(Levy 2004)

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16. Haplodrassus nabozhenkoi Ponomarev, 2023
speciesimage

male Pedipalp, prolateral view
(Ponomarev & Shmatko 2023b)

 speciesimage

male Pedipalp, retrolateral view
(Ponomarev & Shmatko 2023b)

 speciesimage

male Pedipalp, ventral view
(Ponomarev & Shmatko 2023b)
speciesimage

male Pedipalp, ventral view
(Ponomarev & Shmatko 2023b)

 speciesimage

female Epigyne
(Ponomarev & Shmatko 2023b)

 speciesimage

female Vulva, dorsal view
(Ponomarev & Shmatko 2023b)
speciesimage

female Vulva, ventral view
(Ponomarev & Shmatko 2023b)

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17. Haplodrassus nigroscriptus (Simon, 1909)
speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, dorsal view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)
speciesimage

female Pedipalp, lateral view
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Bosmans et al. 2018)
speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

 speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)

 speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

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cp

18. Haplodrassus omissus (O. Pickard-Cambridge, 1872)
speciesimage

male Pedipalp, ventral view
(Bosmans & Van Keer 2012)

 speciesimage

male Pedipalp, ventral view
(Bosmans & Van Keer 2012)

 speciesimage

male Pedipalp, prolateral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)
speciesimage

male Apical part of bulbus
(Denis 1956a)

 speciesimage

male Embolic division, ventral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, dorsal view
(Denis 1956a)
speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)

 speciesimage

male Vulva, ventral view
(Bosmans et al. 2018)
speciesimage

female Epigyne
(Chatzaki et al. 2002a)

 speciesimage

female Epigyne, ventral view
(Denis 1956a)

 speciesimage

female Epigyne
(Bosmans et al. 2018)
speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Vulva
(Chatzaki et al. 2002a)

 speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)
speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)

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19. Haplodrassus orientalis (L. Koch, 1866)
speciesimage

male Pedipalp, retrolateral view
(Kovblyuk et al. 2012b)

 speciesimage

male Pedipalp, ventral view
(Kovblyuk et al. 2012b)

 speciesimage

male Pedipalp, ventral view
(Ponomarev & Tsvetkov 2006)
speciesimage

male Pedipalp, retrolateral view
(Ponomarev & Tsvetkov 2006)

 speciesimage

male Pedipalp, dorsal view
(Naumova et al. 2021)

 speciesimage

male Pedipalp, prolateral view
(Naumova et al. 2021)
speciesimage

male Pedipalp, retrolateral view
(Naumova et al. 2021)

 speciesimage

male Pedipalp, ventral view
(Naumova et al. 2021)

 speciesimage

male Pedipalp, retrolateral view
(Seyyar et al. 2022a)
speciesimage

male Pedipalp, prolateral view
(Seyyar et al. 2022a)

 speciesimage

male Pedipalp, dorso-retrolateral view
(Seyyar et al. 2022a)

 speciesimage

male Pedipalp, ventral view
(Seyyar et al. 2022a)
speciesimage

male Pedipalp, ventral view
(Seyyar et al. 2022a)

 speciesimage

male Bulbus, apical view
(Kovblyuk et al. 2012b)

 speciesimage

male Tip of pedipalp, ventral view
(Esyunin & Tuneva 2020)
speciesimage

female Pedipalp
(Bosmans et al. 2018)

 speciesimage

female Epigyne, ventral view
(Kovblyuk et al. 2012b)

 speciesimage

female Epigyne, ventral view
(Ponomarev & Tsvetkov 2006)
speciesimage

female Epigyne
(L. Koch 1866)

 speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Naumova et al. 2021)
speciesimage

female Epigyne
(Seropian et al. 2023b)

 speciesimage

female Vulva, dorsal view
(Kovblyuk et al. 2012b)

 speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)
speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)

 speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)

 speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)
speciesimage

female Vulva, dorsal view
(Naumova et al. 2021)

 speciesimage

female Vulva, dorsal view
(Seropian et al. 2023b)

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cp

20. Haplodrassus ovatus Bosmans & Hervé, 2018
speciesimage

male Pedipalp, prolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Embolic apophyis, ventral view
(Bosmans et al. 2018)

 speciesimage

male Embolic apophyis, ventral view
(Bosmans et al. 2018)
speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Bosmans et al. 2018)
speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)

 speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)
speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

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cp

21. Haplodrassus ovtchinnikovi Ponomarev, 2008
speciesimage

male Pedipalp, retrolateral view
(Kovblyuk et al. 2009)

 speciesimage

male Pedipalp, ventral view
(Kovblyuk et al. 2009)

 speciesimage

male Palpal tibia, retrolateral view
(Kovblyuk et al. 2009)
speciesimage

female Epigyne, ventral view
(Kovblyuk et al. 2009)

 speciesimage

female Vulva, dorsal view
(Kovblyuk et al. 2009)

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22. Haplodrassus ponomarevi Kovblyuk & Seyyar, 2009
speciesimage

male Pedipalp, retrolateral view
(Kovblyuk et al. 2009)

 speciesimage

male Pedipalp, ventral view
(Kovblyuk et al. 2009)

 speciesimage

male Pedipalp, prolateral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, retrolateral view
(Kovblyuk et al. 2009)
speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)

 speciesimage

female Epigyne, ventral view
(Kovblyuk et al. 2009)

 speciesimage

female Vulva, dorsal view
(Kovblyuk et al. 2009)

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cp

23. Haplodrassus pseudosignifer Marusik, Hippa & Koponen, 1996
speciesimage

male Pedipalp, ventral view
(Kovblyuk et al. 2012b)

 speciesimage

male Bulbus, apical view
(Kovblyuk et al. 2012b)

 speciesimage

male Bulbus, dorsal view
(Šestáková et al. 2022)
speciesimage

male Bulbus, retrolateral view
(Šestáková et al. 2022)

 speciesimage

male Bulbus, ventro-prolateral view
(Šestáková et al. 2022)

 speciesimage

male Tibial apophysis, retrolateral view
(Kovblyuk et al. 2012b)
speciesimage

female Epigyne, ventral view
(Kovblyuk et al. 2012b)

 speciesimage

female Vulva, dorsal view
(Kovblyuk et al. 2012b)

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24. Haplodrassus pugnans (Simon, 1880)
speciesimage

male Pedipalp, ventral view
(Kovblyuk et al. 2012b)

 speciesimage

male Pedipalp, ventral view
(Levy 2004)

 speciesimage

male Bulbus, apical view
(Kovblyuk et al. 2012b)
speciesimage

male Palpal tibia, retrolateral view
(Levy 2004)

 speciesimage

male Tibial apophysis, retrolateral view
(Kovblyuk et al. 2012b)

 speciesimage

female Epigyne, ventral view
(Kovblyuk et al. 2012b)
speciesimage

female Epigyne
(Levy 2004)

 speciesimage

female Vulva, dorsal view
(Kovblyuk et al. 2012b)

 speciesimage

female Vulva, dorsal view
(Levy 2004)

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25. Haplodrassus rhodanicus (Simon, 1914)
speciesimage

male Pedipalp, prolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, retrolateral view
(Bosmans et al. 2018)
speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)

 speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)
speciesimage

female Vulva
(Bosmans et al. 2018)

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26. Haplodrassus rufipes (Lucas, 1846)
speciesimage

male Pedipalp, ventral view
(Simon 1914)

 speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp without embolic part, ventral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Breitling 2020)

 speciesimage

male Embolic division, ventral view
(Bosmans et al. 2018)
speciesimage

male Palpal tibia, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, retrolateral view
(Breitling 2020)
speciesimage

female Epigyne
(Simon 1914)

 speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Bosmans et al. 2018)
speciesimage

female Epigyne, variation
(Simon 1914)

 speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)

 speciesimage

female Vulva
(Bosmans et al. 2018)

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cp

27. Haplodrassus securifer Bosmans & Abrous, 2018
speciesimage

male Pedipalp, prolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Embolic apophyis, ventral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, retrolateral view
(Bosmans et al. 2018)
speciesimage

male Palpal tibia, retrolateral view
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)
speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

 speciesimage

female Vulva
(Bosmans et al. 2018)

 speciesimage

-- Caption missing --
(Bosmans et al. 2018)

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cp

28. Haplodrassus stuxbergi (L. Koch, 1879)
speciesimage

male Pedipalp, ventral view
(Marusik & Logunov 1995b)

 speciesimage

male Embolus, apical view
(Marusik & Logunov 1995b)

 speciesimage

male Palpal tibia, retro-apical view
(Marusik & Logunov 1995b)
speciesimage

female Epigyne, ventral view
(Marusik & Logunov 1995b)

 speciesimage

female Vulva, dorsal view
(Marusik & Logunov 1995b)

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cp

29. Haplodrassus triangularis Bosmans, 2018
speciesimage

male Pedipalp, prolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)
speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)
speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

 speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

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cp

30. Haplodrassus typhon (Simon, 1878)
speciesimage

male Pedipalp, prolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, retrolateral view
(Bosmans et al. 2018)

 speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)
speciesimage

male Pedipalp, ventral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)

 speciesimage

male Palpal tibia, dorso-lateral view
(Bosmans et al. 2018)
speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Epigyne
(Bosmans et al. 2018)

 speciesimage

female Vulva, dorsal view
(Bosmans et al. 2018)
speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

 speciesimage

female Vulva, ventral view
(Bosmans et al. 2018)

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cp

31. Haplodrassus umbratilis gothicus Lohmander, 1942
speciesimage

-- Caption missing --
(WSC 2025)