Gnaphosidae  Pocock, 1898 / Haplodrassus  Chamberlin, 1922   44 species

Haplodrassus signifer  (C. L. Koch, 1839)

Description

Male palp with distinctly keeled distal bulbal apophysis. Prosoma red-brown, eye region darker. Chelicerae darker than prosoma. Legs brighter than prosoma. Opisthosoma provided with copper-coloured hairs.

Body length male: 4.7-7.4 mm
Body length female: 5.1-11.8 mm
Additional information

On dry meadows, on sand and stony debris, lawns, heathland, open forests and damp areas, up to 3000 m in the Alps, in litter and under stones during the day

Frequency: Very frequent, widely distributed

Distribution
  Presence
  No data
Global distribution (WSC 2020): North America, Europe, North Africa, Turkey, Caucasus, Russia (Europe to Far East), Israel, Iran, Central Asia, China, Korea
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List
Albania   ( Deltshev et al., 2011b; ) ||| Algeria   ( Bosmans, unpubl.; ) ||| Austria   ( no references ) ||| Azerbaijan   ( Otto, 2020; ) ||| Belarus   ( no references ) ||| Belgium   ( no references ) ||| Bulgaria   ( Blagoev et al., 2018; ) ||| Croatia   ( no references ) ||| Cyprus   ( Bosmans et al., 2019c; ) ||| Czechia   ( no references ) ||| Denmark   ( no references ) ||| Estonia   ( no references ) ||| Faroe Islands   ( Lissner et al., 2016b; ) ||| Finland   ( Koponen et al., 2016; ) ||| France   ( no references ) ||| France / Corsica   ( no references ) ||| Georgia   ( Otto, 2020; ) ||| Germany   ( no references ) ||| Greece   ( no references ) ||| Greece / Crete   ( no references ) ||| Hungary   ( no references ) ||| Iceland   ( no references ) ||| Ireland   ( no references ) ||| Italy   ( Pantini & Isaia, 2019; ) ||| Italy / Sardinia   ( Pantini & Isaia, 2019; ) ||| Italy / Sicily   ( Pantini & Isaia, 2019; ) ||| Kosovo   ( no references ) ||| Latvia   ( Cera, 2018; ) ||| Liechtenstein   ( no references ) ||| Lithuania   ( no references ) ||| Luxembourg   ( Hermann, 1998; ) ||| Malta   ( Pfliegler et al., 2017; ) ||| Moldova   ( no references ) ||| Montenegro   ( no references ) ||| Morocco   ( Bosmans, unpubl.; ) ||| Netherlands   ( no references ) ||| North Macedonia   ( no references ) ||| Northern Ireland   ( no references ) ||| Norway   ( no references ) ||| Poland   ( no references ) ||| Portugal   ( no references ) ||| Romania   ( no references ) ||| Russia, Central   ( no references ) ||| Russia, Eastern   ( no references ) ||| Russia, Kaliningrad Region   ( no references ) ||| Russia, Northern   ( no references ) ||| Russia, Southern   ( Otto, 2020; ) ||| Russia, Western   ( no references ) ||| Serbia   ( no references ) ||| Slovakia   ( no references ) ||| Slovenia   ( Kostanjšek & Kuntner, 2015; ) ||| Spain   ( no references ) ||| Sweden   ( no references ) ||| Switzerland   ( no references ) ||| Tunisia   ( Bosmans, unpubl.; ) ||| Turkey (Asia)   ( Danışman et al., 2019; ) ||| Turkey (Europe)   ( Danışman et al., 2019; ) ||| Ukraine   ( Polchaninova & Prokopenko, 2019; ) ||| United Kingdom   ( no references ) |||
References

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018)

Bosmans R, Kherbouche-Abrous O, Benhalima S, Hervé C (2018) The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the Maghreb in particular (Araneae: Gnaphosidae) Zootaxa 4451: 1-67 pdf

Bosmans R, Van Keer J, Russell-Smith A, Hadjiconstantis M, Komnenov M, Bosselaers J, Huber S, McCowan D, Snazell R, Decae A, Zoumides C, Kielhorn K-H, Oger P (2019c) Spiders of Cyprus (Araneae). A catalogue of all currently known species from Cyprus. Newsletter of the Belgian Arachnological Society 34, Suppl. 1: 1-173 pdf

Bosmans R. (unpubl.) Provisional list of spiders of North Africa. Database excerpt Aug. 2019

Brændegaard J (1966) Edderkopper: Eller Spindlere I Danmarks Fauna 72: 1-224 pdf

Cera I (2018) The checklist of Latvian spiders (Arachnida: Araneae). Environmental and Experimental Biology 16: 139-152 pdf

Danışman T, Kunt K B, Özkütük R S (2019) The checklist of the spiders of Turkey. Version 2019 [last updated 1 June 2019], online at http://www.spidersofturkey.info

Deltshev C, Vrenozi B, Blagoev G, Lazarov S (2011b) Spiders of Albania – faunistic and zoogeographical review (Arachnida: Araneae). Acta Zoologica Bulgarica 63: 125-144

Helsdingen P J van (2010a) Araneae. In: Fauna Europaea Database (Version 2010.1), online at www.european-arachnology.org

Hermann E (1998) Die Spinnen (Araneae) ausgewählter Halbtrockenrasen im Osten Luxemburgs. Bulletin de la Société des naturalistes Luxembourgeois 99: 189-199 pdf

Koponen S, Fritzén N R, Pajunen T (2016) Checklist of spiders in Finland (Araneae). 6th version, December 2016, online at http://biolcoll.utu.fi/arach/checklist_of_spiders_in_Finland.htm

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Kovblyuk M M, Kastrygina Z A, Omelko M M (2012b) A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae,  Gnaphosidae). Zookeys 205: 59-89 pdf

Lissner J, Jensen J-K, Hansen L J, Simonsen W, Kelduni R, Nissen K (2016b) An updated checklist of spiders (Araneae) of the Faroe Islands. Norwegian Journal of Entomology 63: 197-240 pdf

Locket G H, Millidge A F (1951) British Spiders 1. Ray Soc London

Løvbrekke H (unpubl.)

Noordam A P (1992) Gnaphosidae (Araneae) of the Netherlands, drawings, some notes, and some thoughts about ant-mimicry. Published by the author.

Otto S (2020) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 03.2020. Internet: caucasus-spiders.info.

Pantini P, Isaia M (2019) Araneae.it: the online Catalog of Italian spiders, with addenda on other Arachnid Orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). http://araneae.it. Fragmenta Entomologica 51: 127-152 pdf

Pfliegler W A, Schönhofer A, Niedbała W, Vella P, Sciberras A, Vella A (2017) New records of mites (Acari) and harvestmen (Opiliones) from Malta with a preliminary checklist of Maltese Arachnida. Soil Organisms 89: 85-110 pdf

Platnick N I, Shadab M U (1975) A revision of the spider genera Haplodrassus and Orodrassus (Araneae, Gnaphosidae) in North America. American Museum Novitates 2583: 1-40

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Roberts M J (1995) Spiders of Britain and Northern Europe. Collins Field Guide Bath

Tullgren A (1946) Svensk spindelfauna. 3. Egentiliga spinddlar. Araneae Fam 5-7. Clubionidae, Zoridae och Gnaphosidae. pl. I - XXI Stockholm (Entomol Fören): 1-141

World Spider Catalog (2020) World Spider Catalog. Version 21.0. Natural History Museum Bern, online at http://wsc.nmbe.ch (20.03.2020) doi: 10.24436/2

Updates
17-03-2020 Distribution insert Detail
02-10-2018 Image insert
10-09-2018 Image insert
09-08-2018 Image insert
02-02-2016 Image insert
10-12-2013 Distribution update Detail
01-12-2013 Distribution update Detail