Gnaphosidae  Banks, 1892 / Haplodrassus  Chamberlin, 1922   43 species

Haplodrassus moderatus (Kulczyński, 1897)

Description

Tibial apophysis small, reaches about one third of bulbus length. Epigyne with posteriorly converging margins. Prosoma brown to red-brown, eye region darker. Chelicerae darker than body colour. Legs brown to red-brown, distal leg segments darker. Opisthosoma brown to red-brown.

Body length male: 4.5-6.4 mm
Body length female: 6.5-8.5 mm
Additional information

in humid meadows, moors and bog forests

Distribution
  Presence
  No data
Global distribution (WSC 2026): Europe, Russia (Europe to Far East), China
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List

"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.

Austria   (Milasowszky & Waitzbauer, 2008) ||| Belarus   (Ivanov, 2013) ||| Belgium   (Bosmans & Van Keer, 2017) ||| Czechia   (Růžička & Řezáč, 2022a) ||| Denmark   (Scharff & Gudik-Sørensen, 2006) ||| Estonia   (Vilbaste, 1987) ||| Finland   (Koponen et al., 2016) ||| Germany   (Blick et al., 2016) ||| Hungary   (Samu & Szinetár, 1999) ||| Latvia   (Cera, 2018) ||| Lithuania   (Biteniekytė & Rėlys, 2012) ||| Norway   (Aakra & Hauge, 2003) ||| Poland   (Rozwałka & Stanska, 2008) ||| Romania   (Moscaliuc, 2013) ||| Russia, Eastern   (Tuneva & Esyunin, 2003a) ||| Russia, Northern   (Esyunin & Efimik, 1996a) ||| Russia, Southern   (Otto, 2022; Ponomarev, 2022) ||| Russia, Western   (no references) ||| Serbia   (Grbić et al., 2021) ||| Slovakia   (Gajdoš et al., 1999) ||| Sweden   (Kronestedt, 2001) ||| Switzerland   (Maurer & Hänggi, 1990) ||| Ukraine   (Polchaninova & Prokopenko, 2019) |||
References

Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129 pdf

Almquist S (2007) Swedish Araneae, part 2 – families Dictynidae to Salticidae. Insect Systematics & Evolution, Supplement 63(2006): 285-601 pdf

Biteniekytė M, Rėlys V (2012) The checklist of Lithuanian spiders (Arachnida: Araneae). Biologija 57: 148-158 pdf

Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510 pdf

Bosmans R, Van Keer K (2017) Een herziene soortenlijst van de Belgische spinnen (Araneae). Nieuwsbrief van de Belgische Arachnologische Vereniging 32: 39-69 pdf

Cera I (2018) The checklist of Latvian spiders (Arachnida: Araneae). Environmental and Experimental Biology 16: 139-152 pdf

Esyunin S L, Efimik V E (1996a) Catalogue of the spiders (Arachnida, Aranei) of the Urals. KMK Scientific Press, Moscow, 228 pp. pdf

Fjellberg A (unpubl.) .

Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp.

Grbić G, Hänggi A, Krnjajić S (2021) Spiders (Araneae) of Subotica Sandland (Serbia): additional arguments in environmental protection. Acta Zoologica Academiae Scientiarum Hungaricae 67: 15-61 pdf

Grimm U (1985) Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF) 26: 1-318 pdf

Ivanov V V (2013) The checklist of Belarusian spiders (Arachnida, Araneae). Zoology and Ecology 4: 293-311 pdf

Koponen S, Fritzén N R, Pajunen T (2016) Checklist of spiders in Finland (Araneae). 6th version, December 2016, online at http://biolcoll.utu.fi/arach/checklist_of_spiders_in_Finland.htm pdf

Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001] pdf

Løvbrekke H (unpubl.)

Maurer R, Hänggi A (1990) Katalog der schweizerischen Spinnen. Documenta Faunistica Helvetiae 12 pdf

Milasowszky N, Waitzbauer W (2008) Die Spinnenfauna (Araneae) des Seedammes im Nationalpark Neusiedler See - Seewinkel. Abhandlungen der Zoologisch-Botanischen Gesellschaft Österreich 37: 93-105 pdf

Miller F (1971) Pavouci-Araneida. Klíč zvířeny ČSSR 4: 51-306 pdf

Moscaliuc L A (2013) Contributions to the knowledge of Romanian spider fauna. Steps towards an updated checklist. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 56: 135-142 pdf

Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Roberts M J (1998) Spinnengids. Tirion, Baarn, Netherlands, 397 pp. pdf

Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Scharff N, Gudik-Sørensen O (2006) Katalog over Danmarks edderkopper (Araneae). Catalogue of the spiders of Denmark (Araneae). Entomologiske Meddelelser 74: 3-71 pdf

Tullgren A (1946) Svensk spindelfauna. 3. Egentiliga spinddlar. Araneae Fam 5-7. Clubionidae, Zoridae och Gnaphosidae. Stockholm (Entomol Fören.): 141 pp. pdf

Tuneva T K, Esyunin S L (2003a) A review of the Gnaphosidae fauna of the Urals (Aranei), 3. New species and new records, chiefly from the south Urals. Arthropoda Selecta 11: 223-234 pdf

Vilbaste A (1987) Eesti Ämblikud (Aranei). Annoteeritud nimestik. Estonian spiders (Aranei). An annotated check list. Valgus, Tallinn. 113 pp. & 512 maps on 67 unpag. pages pdf

WSC (2025) World Spider Catalog. Version 26. Natural History Museum Bern, online at http://wsc.nmbe.ch (28.2.2025) doi: 10.24436/2 pdf

Updates
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