Philodromidae: Thanatus C. L. Koch, 1837

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1

Femur/leg II longer than I and IV, embolus tip long and filiform, receptacula without transverse seams

 

bungei-group

  2  

-

Femur/leg IV greater or equal to II and I

  3  

2
(1)

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Prosoma width > 2.5 mm, tegular apophysis indistinct, glandular heads (arrow) broadly connected to receptacula

Thanatus bungei (Kulczyński, 1908)

-

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Prosoma width < 2.5 mm, tegular apophysis swollen membranous (arrow), glandular heads connected to receptacula by small stalks

Thanatus oblongiusculus (Lucas, 1846)

3
(1)

Legs elongated (femur IV/prosoma length > 1,7 in males, > 1,3 in females)

  4  

-

Femur IV/prosoma length < 1,5 in males, < 1,3 in females

  5  

4
(3)

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Retrolateral tibial apophysis with thorn-like tip, median septum of epigyne much longer than wide, receptacula with conspicuous contraction

Thanatus fabricii (Audouin, 1826)

-

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Retrolateral tibial apophysis long tapering and slightly undulating, median septum of epigyne as wide as long, receptacula bulbous

Thanatus lesserti (Roewer, 1951)

5
(3)

Tibia of male palp with 2 apophyses (ventral tibial apophysis, retrolateral tibial apophysis), lateral guide pockets of epigyne evenly curved, receptaculum a simple chamber with no distinct compartmentalization and few seams

  6  

-

Tibia of male palp with 1 apophysis (retrolateral tibial apophysis), lateral guide pockets of epigyne arched in posterior half, receptaculum with distinct compartments and seams

 

formicinus- and sabulosus-groups

  15  

6
(5)

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Embolus gradually tapering into fine straight tip, epigyne with anterior guide pockets

Thanatus pictus L. Koch, 1881

-

Embolus base swollen, embolus tip curved, epigyne without anterior guide pockets

 

striatus-group

  7  

7
(6)

Males

  8  

-

Females

  12  

8
(7)

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Retrolateral tibial apophysis with sharp tip pointing dorsad (lateral view)

Thanatus mikhailovi Logunov, 1996

female

-

Retrolateral tibial apophysis in lateral view with rounded tip

  9  

9
(8)

Retrolateral tibial apophysis in lateral view directed dorsad, embolus tip broadly twisted

  10  

-

Retrolateral tibial apophysis in lateral view directed distad/ventrad, embolus tip tightly twisted

  11  

10
(9)

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Embolus gradually narrowing with long tip

Thanatus atratus Simon, 1875

female

-

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Embolus base strongly swollen with apical bulge

Thanatus vulgaris Simon, 1870

female

11
(9)

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Prosoma width > 2 mm, prosoma uniformly dark brown

Thanatus firmetorum Muster & Thaler, 2003

female

-

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Prosoma width < 2 mm, prosoma with light median and lateral bands

Thanatus striatus C. L. Koch, 1845

female

12
(7)

Epigynal fold and lateral guide pockets in close contact

  13  

-

Epigynal fold and lateral guide pockets well separated

  14  

13
(12)

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Lateral guide pockets convex

Thanatus striatus C. L. Koch, 1845

male

-

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Lateral guide pockets concave

Thanatus mikhailovi Logunov, 1996

male

14
(12)

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Median septum wider than long

Thanatus firmetorum Muster & Thaler, 2003

male

-

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Median septum slightly longer than wide

Thanatus atratus Simon, 1875

male

-

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Median septum twice as long as wide

Thanatus vulgaris Simon, 1870

male

15
(5)

Males

  16  

-

Females

  23  

16
(15)

Tip of retrolateral tibial apophysis bifid

  17  

-

Retrolateral tibial apophysis with pointed tip

  19  

17
(16)

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Lobes of the tip of retrolateral tibial apophysis rounded

Thanatus arenarius L. Koch, 1872

female

-

Lobes of the tip of retrolateral tibial apophysis acuminate

  18  

18
(17)

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Cymbium length > 0.8 mm, tegular apophysis long

Thanatus imbecillus L. Koch, 1878

female

-

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Cymbium length < 0.8 mm, tegular apophysis short

Thanatus meronensis Levy, 1977

female

19
(16)

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Retrolateral tibial apophysis slim, much longer than wide

Thanatus sabulosus (Menge, 1875)

female

-

Retrolateral tibial apophysis robust, almost as wide as long

  20  

20
(19)

Retrolateral tibial apophysis keel-shaped, directed retrolaterad

  21  

-

Retrolateral tibial apophysis triangular, directed dorso-distad

  22  

21
(20)

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Tip of retrolateral tibial apophysis not curved, embolus originating at 9 o´clock position on tegulum

Thanatus coloradensis Keyserling, 1880

female

-

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Tip of retrolateral tibial apophysis slightly curved, embolus originating at 10:30 o´clock position on tegulum

Thanatus kitabensis Charitonov, 1946

female

22
(20)

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Tegular apophysis a bulge

Thanatus arcticus Thorell, 1872

female

-

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Tegular apophysis a chitinous blade

Thanatus formicinus (Clerck, 1757)

female

23
(15)

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Median septum round, converging anteriorly

Thanatus arenarius L. Koch, 1872

male

-

Not so

  24  

24
(23)

Lateral guide pockets forming semicircular arches

  25  

-

Lateral guide pockets S-shaped

  26  

25
(24)

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Epigyne wider than long

Thanatus meronensis Levy, 1977

male

-

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Epigyne as wide as long

Thanatus imbecillus L. Koch, 1878

male

26
(24)

Epigyne wider than long, receptacula with a couple of seams

  27  

-

Epigyne longer than wide, receptacula with multiple seams

  28  

27
(26)

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Prosoma width < 2 mm

Thanatus sabulosus (Menge, 1875)

male

-

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Prosoma width > 2 mm

Thanatus arcticus Thorell, 1872

male

28
(26)

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Median septum strongly elongated, lateral guide pockets weakly bulged

Thanatus coloradensis Keyserling, 1880

male

-

Median septum less elongated, lateral guide pockets strongly bulged

  29  

29
(28)

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Medial epigynal pocket present

Thanatus kitabensis Charitonov, 1946

male

-

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Without medial epigynal pocket

Thanatus formicinus (Clerck, 1757)

male

This key is a kind contribution by Christoph Muster.

Further taxa

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1. Thanatus dissimilis Denis, 1960

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2. Thanatus flavescens O. Pickard-Cambridge, 1876

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3. Thanatus flavidus Simon, 1875

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4. Thanatus flavus O. Pickard-Cambridge, 1876

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5. Thanatus fornicatus Simon, 1897

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6. Thanatus fuscipes Denis, 1937

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7. Thanatus fuscipes concolor Denis, 1957

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8. Thanatus lanceolatus Simon, 1875

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9. Thanatus lineatipes Simon, 1870

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10. Thanatus mongolicus (Schenkel, 1936)

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11. Thanatus oblongiusculus atomarius (Simon, 1932)

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12. Thanatus okayi Karol, 1966

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13. Thanatus ornatus (Lucas, 1846)

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14. Thanatus rayi Simon, 1875

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15. Thanatus setiger (O. Pickard-Cambridge, 1872)

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16. Thanatus ubsunurensis Logunov, 1996

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17. Thanatus validus Simon, 1875

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18. Thanatus virgulatipes Wunderlich, 2023

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19. Thanatus vulgaris creticus Kulczyński, 1903