Philodromidae  Thorell, 1870 / Thanatus  C. L. Koch, 1837   35 species

Thanatus atratus  Simon, 1875

Description

Male palp with small, rounded tegular apophysis. Embolus uniformly rounded. Median epigynal plate distinctly separating lateral pockets. Legs grey or light brown and spotted. Opisthosoma grey or light brown.

Body length male: 3.1-4.7 mm
Body length female: 4.6-6.1 mm
Additional information

In dry and semi-dry steppe areas, on sea coasts and salt places, hibernation as juveniles

Distribution
  Presence
  No data
Global distribution (WSC 2020): Europe, Turkey, Caucasus, Russia (Europe to South Siberia), Kazakhstan, Iran, Korea, Japan
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List
Albania   (Vrenozi & Jäger, 2012) ||| Andorra   (Branco et al., 2019) ||| Austria   (no references) ||| Bulgaria   (Blagoev et al., 2018) ||| Croatia   (no references) ||| Czechia   (no references) ||| France   (no references) ||| Georgia   (Otto, 2020) ||| Germany   (no references) ||| Greece   (no references) ||| Greece / Crete   (Bosmans et al., 2013) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Pantini & Isaia, 2019) ||| North Macedonia   (no references) ||| Norway   (Aakra & Hauge, 2003) ||| Poland   (no references) ||| Portugal   (Branco et al., 2019) ||| Romania   (no references) ||| Russia, Eastern   (no references) ||| Russia, Southern   (Otto, 2020) ||| Slovenia   (Kostanjšek & Kuntner, 2015) ||| Spain   (Branco et al., 2019) ||| Sweden   (Kronestedt, 2001) ||| Switzerland   (no references) ||| Turkey (Asia)   (Danışman et al., 2019) ||| Ukraine   (Polchaninova & Prokopenko, 2019) |||
References

Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129 pdf

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Bosmans R, Van Keer J, Russell-Smith A, Kronestedt T, Alderweireldt M, Bosselaers J, De Koninck H (2013) Spiders of Crete (Araneae). A catalogue of all currently known species from the Greek island of Crete. Nieuwsbrief van de Belgische Arachnologische Vereniging 28 (Suppl. 1): 1-147 pdf

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). [N.B.: few records from this paper are not taken over and not commented] Zootaxa 4614: 201-254 pdf

Danışman T, Kunt K B, Özkütük R S (2019) The checklist of the spiders of Turkey. Version 2019 [last updated 1 June 2019], online at http://www.spidersofturkey.info pdf

Hansen H (1995) Über die Arachniden-Fauna von urbanen Lebensräumen in Venedig - III. Die epigäischen Spinnen eines Stadtparkes (Arachnida: Araneae). Bollettino del Museo Civico di Storia Naturale di Venezia 44: 7-36 pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Kronestedt T (1983) Spiders (Araneae) on the Great Alvar of the island of Öland, S Sweden. Entomologisk Tidskrift 104: 183-212 pdf

Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001] pdf

Logunov D V (1996b) A critical review of the spider genera Apollophanes O. P.-Cambridge, 1898 and Thanatus C. L. Koch, 1837 in North Asia (Araneae, Philodromidae). Revue Arachnologique 11: 133-202 pdf

Muster C, Thaler K (2003) The Thanatus striatus species group in the eastern Alps, with description of Thanatus firmetorum sp. n. (Araneae: Philodromidae). Bulletin of the British Arachnological Society 12: 376-382 pdf

Otto S (2020) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 03.2020. Internet: caucasus-spiders.info.

Pantini P, Isaia M (2019) Araneae.it: the online catalog of Italian spiders, with addenda on other arachnid orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). http://araneae.it. Fragmenta Entomologica 51: 127-152 pdf

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Pozzi S, Hänggi A (1998) Araignées nouvelles ou peu connues de la Suisse (Arachnida: Araneae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 71: 33-47 pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Szita É, Samu F (2000) Taxonomical review of Thanatus species (Philodromidae, Araneae) of Hungary. Acta Zoologica Academiae Scientiarum Hungaricae 46: 155-179 pdf

Van Keer J, Van Keer K, De Koninck H, Ramel G (2010) The arachnofauna (Araneae) of wetland Kerkini (Macedonia-Northern Central Greece). Acta Zoologica Bulgarica 62: 141-160 pdf

Vrenozi B, Jäger P (2012) A faunistic study on ground-dwelling spiders (Araneae) in the Tirana district, Albania. Arachnologische Mitteilungen 44: 81-87 pdf

World Spider Catalog (2020) World Spider Catalog. Version 21.0. Natural History Museum Bern, online at http://wsc.nmbe.ch (20.03.2020) doi: 10.24436/2 pdf

Updates
26-07-2020 Image insert
24-09-2015 Distribution update Detail
12-11-2013 Image insert
15-10-2013 Distribution update Detail