Clubionidae: Clubiona Latreille, 1804

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1
Opisthosoma also in posterior half with distinct pattern consisting of a median band, chevrons or arcuate stripes

  3  

- Opisthosoma only in anterior half with wedge-shaped cardiac mark or without pattern

  2  

2
(1)

Chelicerae distinctly darker (mostly dark brown to black) than prosoma (mostly light to middle brown) -> see also C. lutescens

  12  

- Chelicerae not darker than prosoma

  28  

3
(1)

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Bulbus simple, unstructured and large; epigynal groove 1 1/2 of its length apart from epigastric furrow

Clubiona corticalis (Walckenaer, 1802)

- Different

  4  

4
(3)
Ventral tibial apophysis arrow-shaped, with barb; receptaculum seminis not distinctly visible through anterior half of epigyne; epigyne wider than long

  5  

- Tibial apophysis not arrow-shaped, without barb; epigyne at least as long as wide; 2 (mostly) circular receptacula seminis visible through anterior half of epigyne

  6  

5
(4)

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Tibial apophysis blunt and with a barb; posterior epigynal margin with distinct quadrangular lobe

Clubiona kulczynskii Lessert, 1905

-

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Tibial apophysis pointed and with 2 barbs; epigyne as fig.

Clubiona subsultans Thorell, 1875

6
(4)
Male pedipalp with simple undivided tibial apophysis; epigyne with large, ± heart-shaped groove

  7  

- Tibial apophysis tapering into a tip, narrowed at base or slightly bent; epigyne without distinct groove

  9  

7
(6)

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Tibial apophysis and tibia approx. equal in length; epigynal groove roundish trapezoid

Clubiona juvenis Simon, 1878

- Tibial apophysis smaller than half the length of tibia; epigyne different

  8  

8
(7)

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Tibial apophysis rounded; epigynal groove heart-shaped; entrance ducts spiral; chelicerae hardly swollen

Porrhoclubiona genevensis (L. Koch, 1866)

-

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Tibial apophysis pointed; epigynal groove as fig.; chelicerae strikingly swollen

Porrhoclubiona vegeta (Simon, 1918)

9
(6)

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Ventral tibial apophysis with long, pointed, bent process; epigynal groove with semicircular entrance ducts, visible through epigyne

Clubiona comta C. L. Koch, 1839

- Different

  10  

10
(9)

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Tibial apophysis approx. as long as wide; median part of posterior epigynal margin strongly sclerotised

Clubiona alpicola Kulczyński, 1882

- Tibial apophysis distinctly longer than wide; epigyne different

  11  

11
(10)

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Embolus short, bent; tibial apophysis as fig.; posterior epigynal margin backwardly protruding

Clubiona hilaris Simon, 1878

-

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Embolus long, straight; Tibial apophysis as fig.; posterior epigynal margin deeply indented with distinct, protruding median part which can be longish rectangular to broad triangular

Clubiona saxatilis L. Koch, 1867

12
(2)

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Tibial apophysis with 4 processes; tibia much larger than bulbus; epigyne beak-like elongated

Clubiona caerulescens L. Koch, 1867

- Different

  13  

13
(12)
Males

  14  

- Females

  21  

14
(13)
Ventral tibial apophysis pointed, arrow-shaped, with barb

  15  

- Tibial apophysis different

  17  

15
(14)

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Ventral tibial apophysis with 2 pointed and one round process

Clubiona stagnatilis Kulczyński, 1897

female

- Different

  16  

16
(15)

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Dorsal tibial apophysis broad and round

Clubiona rosserae Locket, 1953

female

-

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Dorsal tibial apophysis pointed

Clubiona norvegica Strand, 1900

female

17
(14)
Tibia next to broadly rounded apophysis with U-like projecting, narrow apophysis

  18  

-

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Different; tibial apophysis as fig.

Clubiona phragmitis C. L. Koch, 1843

female

18
(17)

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U-like projecting branch of tibial apophysis broadened and indented at tip

Clubiona similis L. Koch, 1867

female

- Different

  19  

19
(18)

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Narrow branch of tibial apophysis rounded at tip

Clubiona terrestris Westring, 1851

female

- Different

  20  

20
(19)

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Narrow branch of tibial apophysis slightly grooved at tip; chelicerae approx. half as long as prosoma

Clubiona neglecta O. Pickard-Cambridge, 1862

female

-

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Narrow branch of tibial apophysis slightly broadened at tip; chelicerae approx. as long as prosoma

Clubiona pseudoneglecta Wunderlich, 1994

female

21
(13)
Posterior epigynal margin with medially protruding lobe

  22  

- Different

  23  

22
(21)

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Epigyne and vulva as figs.

Clubiona neglecta O. Pickard-Cambridge, 1862

male

-

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Epigyne similar, but receptacula seminis more longish than in neglecta

Clubiona pseudoneglecta Wunderlich, 1994

male

23
(21)
Epigyne wider than long, posterior margin more strongly sclerotised

  24  

- Epigyne longer than wide

  26  

24
(23)

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2 round receptacula seminis in median epigynal region, distinctly visible

Clubiona rosserae Locket, 1953

male

- Different

  25  

25
(24)

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Receptacula seminis in anterior epigynal region, visible through epigyne

Clubiona stagnatilis Kulczyński, 1897

male

-

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Receptacula seminis in posterior epigynal region, visible through epigyne

Clubiona norvegica Strand, 1900

male

26
(23)

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Entrance duct distinctly visible, obtuse-angled

Clubiona terrestris Westring, 1851

male

- Different

  27  

27
(26)

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Epigyne as fig.; copulatory opening more slit-like than round (different as in drawings by Wiehle)

Clubiona similis L. Koch, 1867

male

-

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Epigyne and vulva as fig.

Clubiona phragmitis C. L. Koch, 1843

male

28
(2)
Males

  29  

- Females

  38  

29
(28)

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Tibial apophysis spear-shaped

Clubiona reclusa O. Pickard-Cambridge, 1863

female

- Different

  30  

30
(29)
Tibia with a dorsal and a ventral apophysis, both ± equal in width and blunt

  31  

- Different

  32  

31
(30)

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Ventral tibial apophysis half as long as dorsal tibial apophysis

Clubiona lutescens Westring, 1851

female

-

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Both tibial apophysis ± equal in length; ventral tibial apophysis slightly stalked, slightly varying in form

Clubiona frutetorum L. Koch, 1867

female

32
(30)
Tibial apophysis with a unstructured dorsal apophysis, at least as long as tibia, ± as wide as tibia at base

  33  

- Different

  36  

33
(32)
Tibial apophysis broad, ± blunt

  34  

- Tibial apophysis cut at an angle, ± pointed

  35  

34
(33)

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Tibial apophysis ± as long as tibia

Clubiona trivialis C. L. Koch, 1843

female

-

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Tibial apophysis twice as long as tibia

Clubiona marmorata L. Koch, 1866

female

35
(33)

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Tibial apophysis with distinctly stepped tip

Clubiona subtilis L. Koch, 1867

female

-

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Tibial apophysis uniformly tapering distally

Clubiona diversa O. Pickard-Cambridge, 1862

female

36
(32)

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Tibia with a blunt dorsal and a pointed ventral apophysis

Clubiona germanica Thorell, 1871

female

- Different

  37  

37
(36)

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Dorsal tibial apophysis with thin, bent process, forked at tip

Clubiona brevipes Blackwall, 1841

female

-

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Tibial apophysis different

Clubiona pallidula (Clerck, 1757)

female

38
(28)

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Entrance ducts visible as arcuate slits

Clubiona lutescens Westring, 1851

male

- Different

  39  

39
(38)

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Entrance ducts slit-shaped, diverging posteriorly

Clubiona marmorata L. Koch, 1866

male

- Different

  40  

40
(39)
Epigynal groove distinctly bordered posteriorly

  41  

- Different

  45  

41
(40)
Epigynal groove undivided

  43  

- Epigynal groove bipartite

  42  

42
(41)

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Epigynal groove broadly heart-shaped, on each side with a shallow depression

Clubiona subtilis L. Koch, 1867

male

-

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Epigynal groove without depression

Clubiona germanica Thorell, 1871

male

43
(41)

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Entrance ducts semicircular, visible at posterior epigynal margin

Clubiona brevipes Blackwall, 1841

male

- Different

  44  

44
(43)

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Epigynal groove round; epigynal region with trapezoid chitinous plate, only a few structures visible through this plate

Clubiona trivialis C. L. Koch, 1843

male

-

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Epigynal groove longish oval

Clubiona diversa O. Pickard-Cambridge, 1862

male

45
(40)

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Posterior epigynal margin nose-like produced posteriorly

Clubiona reclusa O. Pickard-Cambridge, 1863

male

- Different

  46  

46
(45)

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Epigyne as fig.; inner receptacula seminis larger than outer atria

Clubiona pallidula (Clerck, 1757)

male

-

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Epigyne with roundish imprint of receptacula seminis, marked off anteriorly by distinctly stronger sclerotisation

Clubiona frutetorum L. Koch, 1867

male

Further taxa

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1. Clubiona abboti L. Koch, 1866

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2. Clubiona aducta Simon, 1932

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3. Clubiona alexeevi Mikhailov, 1990

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4. Clubiona andreinii Caporiacco, 1936

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5. Clubiona bashkirica Mikhailov, 1992

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6. Clubiona caucasica Mikhailov & Otto, 2017

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7. Clubiona congentilis Kulczyński, 1913

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8. Clubiona corticalis concolor Kulczyński, 1897

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9. Clubiona corticalis nigra Simon, 1878

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10. Clubiona deterrima Strand, 1904

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11. Clubiona frisia Wunderlich & Schuett, 1995

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12. Clubiona golovatchi Mikhailov, 1990

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13. Clubiona hyrcanica Mikhailov, 1990

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14. Clubiona liachviana Mcheidze, 1997

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15. Clubiona mazandaranica Mikhailov, 2003

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16. Clubiona mykolai Mikhailov, 2003

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17. Clubiona pseudosimilis Mikhailov, 1990

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18. Clubiona riparia L. Koch, 1866

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19. Clubiona saltuum Kulczyński, 1898

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20. Clubiona straminea O. Pickard-Cambridge, 1872