Zodariidae: Zodarion Walckenaer, 1826

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1
Males

  2  

- Females

  17  

2
(1)
Tibial apophysis slender, in lateral view more than twice as long as diameter of tibia

  3  

- Tibial apophysis robust, less than twice as long as diameter of tibia

  5  

3
(2)

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Tibial apophysis terminally with 2 pointed teeth

Zodarion pseudoelegans Denis, 1934

female

- Different

  4  

4
(3)

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Distal part of tibial apophysis broad, with a pointed tooth

Zodarion elegans (Simon, 1873)

female

-

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Distal part of tibial apophysis narrower, without tooth

Zodarion gracilitibiale Denis, 1934

5
(2)
Tegulum projecting, covers base of retinaculum and/or embolus

  6  

- Tegulum not projecting

  9  

6
(5)

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Tibial apophysis without tooth or hook; femur dark brown; prosoma dark brown; opisthosoma dark violet

Zodarion germanicum (C. L. Koch, 1837)

female

- Tibial apophysis with tooth or hook

  7  

7
(6)

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Tibial apophysis with robust terminal hook; pale coloured species; body length 1.7-2.2 mm

Zodarion pusio Simon, 1914

female

- Different

  8  

8
(7)

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Tibial apophysis longer than diameter of tibia; body length 1.2-1.6 mm

Zodarion emarginatum (Simon, 1873)

female

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Tibial apophysis as long as diameter of tibia; body length 1.5-1.8 mm

Zodarion ruffoi Caporiacco, 1951

female

9
(5)
Tibial apophysis concave subterminally; embolus with second tooth

  10  

- Tibial apophysis not concave subterminally; embolus with one tooth

  13  

10
(9)

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Second embolic tooth very long; prosoma pale red-brown; femora I-II red-brown; femora III-IV and other segments yellow-brown; opisthosoma dark grey to brown; female unknown

Zodarion fulvonigrum (Simon, 1874)

- Different

  11  

11
(10)

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Base of embolus concave; retinaculum longer than wide, basal and distal branches strongly elongated; femur yellowish brown; prosoma light brown with dark spots; opisthosoma brown-violet

Zodarion rubidum Simon, 1914

female

- Base of embolus not concave; retinaculum as long as wide, branches less elongated

  12  

12
(11)

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Embolic tooth approx. equal in length

Zodarion fuscum (Simon, 1870)

female

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Dorsal embolic tooth longer than ventral tooth; prosoma yellowish to orange-brown, eyes and margin darker; legs light yellow to yellow-orange; opisthosoma brown-black

Zodarion couseransense Bosmans, 1997

female

13
(9)

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Tip of tibial apophysis very slender; body length 1.8-2.3 mm; prosoma brown to dark brown with dark pattern; legs yellowish orange; opisthosoma dorsally dark purple-brown, uniformly pale yellow ventrally

Zodarion hamatum Wiehle, 1964

female

- Different

  14  

14
(13)
Distal part of retinaculum broad, less than twice as long as wide

  15  

- Distal part of retinaculum narrow, 3-4 times as long as wide

  16  

15
(14)

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Tip of embolus bent; body length 1.6-2.9 mm; prosoma brown to dark brown with dark pattern; legs yellowish orange; opisthosoma dorsally dark purple-brown, uniformly pale yellow ventrally

Zodarion italicum (Canestrini, 1868)

female

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Tip of embolus straight; body length 2.2-2.9 mm

Zodarion confusum Denis, 1935

female

16
(14)

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Embolus with ridge, relatively robust at tip; body length 2.1-2.8 mm

Zodarion vicinum Denis, 1935

female

-

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Different; femur mostly slightly darkened; prosoma red-brown; opisthosoma brown-violet; body length of males 1.7-2.5 mm

Zodarion gallicum (Simon, 1873)

female

17
(1)
Epigyne with pronounced median groove

  18  

- Epigyne without pronounced median groove

  19  

18
(17)

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Median epigynal groove sausage-shaped as fig.

Zodarion pseudoelegans Denis, 1934

male

-

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Median epigynal groove as fig.; body length 2.8-3.8 mm

Zodarion elegans (Simon, 1873)

male

19
(17)
Posterior epigynal margin with median incision

  20  

- Posterior epigynal margin without median incision

  21  

20
(19)

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Incision angled; femur dark brown; prosoma dark brown; opisthosoma dark violet; body length of females 3-5 mm

Zodarion germanicum (C. L. Koch, 1837)

male

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Incision rounded; body length 1.8-2.0 mm

Zodarion ruffoi Caporiacco, 1951

male

21
(19)

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Epigyne hardly chitinised; pale coloured species; body length 2.6-3.0 mm

Zodarion pusio Simon, 1914

male

- Different

  22  

22
(21)
Anterior margin of posterior median plate of epigyne chitinously tightened

  23  

- Different

  25  

23
(22)

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Posterior epigynal median plate triangular; femur yellowish brown; prosoma light brown, with dark spots; opisthosoma brown-violet; body length of female 3.2-4.5 mm

Zodarion rubidum Simon, 1914

male

- Posterior epigynal median plate trapezoid

  24  

24
(23)

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Epigyne as fig.

Zodarion fuscum (Simon, 1870)

male

-

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Epigyne as fig.; body length 3.8-4.0 mm; prosoma yellowish to orange-brown, eyes and margin darker; legs light yellow to yellow-orange; opisthosoma brown-black

Zodarion couseransense Bosmans, 1997

male

25
(22)
Posterior median plate trapezoid or triangular

  26  

- Posterior median plate square or rectangular

  28  

26
(25)

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Posterior median plate much wider than long, trapezoid, the lateral margins marked with dark stripes; body length 2.3-3.8 mm

Zodarion hamatum Wiehle, 1964

male

- Different

  27  

27
(26)

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Spermathecae with atrium, clearly visible; body length 2.1-4-3 mm

Zodarion italicum (Canestrini, 1868)

male

-

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Different; body length 2.6-4.4 mm

Zodarion confusum Denis, 1935

male

28
(25)

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Posterior median plate narrow, as long as wide, with dark narrow lateral stripes; body length of female 2.5-4.0 mm

Zodarion gallicum (Simon, 1873)

male

- Different

  29  

29
(28)

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Epigyne as fig.; body length 2.7-3.8 mm

Zodarion vicinum Denis, 1935

male

-

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Epigyne as fig.; body length 1.2-1.6 mm

Zodarion emarginatum (Simon, 1873)

male

-

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Male prosoma densely granulated, dark brown, black in eye region and medially; female prosoma yellowish brown; male legs yellow-brown, femora brown; female legs yellow; male opisthosoma shiny black, dorsally granulated, dark purple laterally and ventrally; female opisthosoma uniformly purple-brown, laterally with large light brown spots

Zodarion granulatum Kulczyński, 1908

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Body length of males 2.2 mm

Zodarion graecum (C. L. Koch, 1843)

Further taxa

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1. Zodarion abantense Wunderlich, 1980

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2. Zodarion abnorme Denis, 1952

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3. Zodarion aculeatum Chyzer, 1897

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4. Zodarion affine (Simon, 1870)

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5. Zodarion agricola Bouseksou & Abrous, 2021

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6. Zodarion alacre (Simon, 1870)

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7. Zodarion albipatellare Bosmans, 2009

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8. Zodarion alentejanum Pekár & Carvalho, 2011

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9. Zodarion algarvense Bosmans, 1994

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10. Zodarion algiricum (Lucas, 1846)

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11. Zodarion andalusiacum Jocqué, 1991

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12. Zodarion arabelae Bosmans, 2009

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13. Zodarion arachnaio Bosmans, 2009

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14. Zodarion atlanticum Pekár & Cardoso, 2005

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15. Zodarion attikaense Wunderlich, 1980

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16. Zodarion aurorae Weiss, 1982

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17. Zodarion azrouense Bosmans & Benhalima, 2020

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18. Zodarion bacelarae Pekár, 2003

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19. Zodarion barbarae Bosmans, 2009

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20. Zodarion beroni Komnenov & Chatzaki, 2016

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21. Zodarion beticum Denis, 1957

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22. Zodarion bicoloripes (Denis, 1959)

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23. Zodarion bigaense Bosmans, Özkütük, Varli & Kunt, 2014

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24. Zodarion blagoevi Bosmans, 2009

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25. Zodarion bosmansi Pekár & Cardoso, 2005

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26. Zodarion bozdagense Coşar, 2021

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27. Zodarion brevicephalus Wunderlich, 2023

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28. Zodarion caporiaccoi Roewer, 1942

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29. Zodarion caucasicum Dunin & Nenilin, 1987

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30. Zodarion cesari Pekár, 2011

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31. Zodarion christae Bosmans, 2009

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32. Zodarion costablancae Bosmans, 1994

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33. Zodarion costapratae Pekár, 2011

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34. Zodarion crewsae Coşar, Danışman & Kunt, 2022

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35. Zodarion cyrenaicum Denis, 1935

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36. Zodarion danismani Coşar, 2021

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37. Zodarion deltshevi Bosmans, 2009

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38. Zodarion diatretum Denis, 1935

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39. Zodarion dispar Denis, 1935

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40. Zodarion duriense Cardoso, 2003

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41. Zodarion emilijae Deltshev & Naumova, 2022

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42. Zodarion epirense Brignoli, 1984

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43. Zodarion ericorum Bosmans, 2020

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44. Zodarion evvoia Bosmans, 2009

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45. Zodarion expers (O. Pickard-Cambridge, 1876)

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46. Zodarion fazanicum Denis, 1938

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47. Zodarion frenatum Simon, 1885

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48. Zodarion gaziantepense Danışman & Coşar, 2021

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49. Zodarion gregua Bosmans, 1994

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50. Zodarion guadianense Cardoso, 2003

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51. Zodarion hauseri Brignoli, 1984

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52. Zodarion immaculatum Denis, 1962

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53. Zodarion imroz Dimitrov, 2020

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54. Zodarion isabellinum (Simon, 1870)

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55. Zodarion izmirense Danışman & Coşar, 2020

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56. Zodarion jansseni Bosmans, 2009

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57. Zodarion jeanclaudeledouxi Bosmans & Benhalima, 2020

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58. Zodarion jozefienae Bosmans, 1994

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59. Zodarion kabylianum Denis, 1937

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60. Zodarion karpathos Bosmans, 2009

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61. Zodarion killini Bosmans, 2009

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62. Zodarion konradi Bosmans, 2009

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63. Zodarion korgei Wunderlich, 1980

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64. Zodarion kossamos Bosmans, 2009

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65. Zodarion kunti Coşar, Danışman & Yağmur, 2021

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66. Zodarion ludibundum Simon, 1914

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67. Zodarion lusitanicum Cardoso, 2003

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68. Zodarion lutipes (O. Pickard-Cambridge, 1872)

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69. Zodarion machadoi Denis, 1939

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70. Zodarion maculatum (Simon, 1870)

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71. Zodarion maghrebense Bosmans & Benhalima, 2020

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72. Zodarion mallorca Bosmans, 1994

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73. Zodarion marginiceps Simon, 1914

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74. Zodarion merlijni Bosmans, 1994

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75. Zodarion mesranense Bouragba & Bosmans, 2012

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76. Zodarion messiniense Bosmans, 2009

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77. Zodarion minutum Bosmans, 1994

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78. Zodarion modestum (Simon, 1870)

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79. Zodarion montesacrense Bosmans, 2019

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80. Zodarion morosoides Bosmans, 2009

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81. Zodarion morosum Denis, 1935

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82. Zodarion mostafai Benhalima & Bosmans, 2020

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83. Zodarion murphyorum Bosmans, 1994

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84. Zodarion musarum Brignoli, 1984

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85. Zodarion namrun Dimitrov, 2024

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86. Zodarion nigriceps (Simon, 1873)

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87. Zodarion nitidum (Audouin, 1826)

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88. Zodarion noordami Bosmans, 2009

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89. Zodarion ogeri Bosmans & Benhalima, 2020

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90. Zodarion ohridense Wunderlich, 1973

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91. Zodarion ozkutuki Coşar & Danışman, 2021

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92. Zodarion pacificum Bosmans, 2009

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93. Zodarion pallidum Denis, 1952

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94. Zodarion pantaleonii Bosmans & Pantini, 2019

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95. Zodarion parastyliferum Wunderlich, 2022

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96. Zodarion petrobium Dunin & Zacharjan, 1991

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97. Zodarion pileolonotatum Denis, 1935

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98. Zodarion pirini Drensky, 1921

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99. Zodarion pseudonigriceps Bosmans & Pantini, 2019

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100. Zodarion pythium Denis, 1935

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101. Zodarion remotum Denis, 1935

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102. Zodarion reticulatum Kulczyński, 1908

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103. Zodarion rhodiense Caporiacco, 1948

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104. Zodarion robertbosmans Wunderlich, 2017

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105. Zodarion rudyi Bosmans, 1994

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106. Zodarion samos Bosmans, 2009

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107. Zodarion santorini Bosmans, 2009

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108. Zodarion sardum Bosmans, 1997

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109. Zodarion scutatum Wunderlich, 1980

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110. Zodarion segurense Bosmans, 1994

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111. Zodarion sharurense Zamani & Marusik, 2022

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112. Zodarion siirtense Coşar, 2021

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113. Zodarion soror (Simon, 1873)

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114. Zodarion spinibarbe Wunderlich, 1973

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115. Zodarion styliferum (Simon, 1870)

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116. Zodarion styliferum extraneum Denis, 1935

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117. Zodarion sungar (Jocqué, 1991)

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118. Zodarion talyschicum Dunin & Nenilin, 1987

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119. Zodarion thoni Nosek, 1905

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120. Zodarion timidum (Simon, 1874)

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121. Zodarion trianguliferum Denis, 1952

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122. Zodarion tuber Wunderlich, 2022

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123. Zodarion tunetiacum Strand, 1906

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124. Zodarion turcicum Wunderlich, 1980

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125. Zodarion turkesi Coşar & Danışman, 2021

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126. Zodarion valentii Bosmans, Loverre & Addante, 2019

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127. Zodarion van Bosmans, 2009

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128. Zodarion vanimpei Bosmans, 1994

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129. Zodarion vankeerorum Bosmans, 2009

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130. Zodarion varoli Akpınar, 2016

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131. Zodarion viduum Denis, 1937

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132. Zodarion walsinghami Denis, 1937

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133. Zodarion weissi Deltshev & Naumova, 2022

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134. Zodarion wesolowskae Bosmans & Benhalima, 2020

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135. Zodarion yagmuri Coşar & Danışman, 2021

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136. Zodarion zorba Bosmans, 2009