Philodromidae  Thorell, 1869 / Thanatus  C. L. Koch, 1837   37 species

Thanatus atratus  Simon, 1875

Description

Male palp with small, rounded tegular apophysis. Embolus uniformly rounded. Median epigynal plate distinctly separating lateral pockets. Legs grey or light brown and spotted. Opisthosoma grey or light brown.

Body length male: 3.1-4.7 mm
Body length female: 4.6-6.1 mm
Additional information

In dry and semi-dry steppe areas, on sea coasts and salt places, hibernation as juveniles

Distribution
  Presence
  No data
Global distribution (WSC 2024): Europe, Turkey, Caucasus, Russia (Europe to South Siberia), Kazakhstan, Iran, Korea, Japan
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List

"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.

Albania   (Vrenozi & Jäger, 2012) ||| Andorra   (Branco et al., 2019) ||| Armenia   (Zarikian, 2021) ||| Austria   (no references) ||| Bulgaria   (Blagoev et al., 2018) ||| Croatia   (Grbac et al., 2019) ||| Czechia   (Růžička & Řezáč, 2022a) ||| France   (no references) ||| Georgia   (Otto, 2022) ||| Germany   (no references) ||| Greece   (Van Keer et al., 2010) ||| Greece / Crete   (Bosmans et al., 2013) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Pantini & Isaia, 2019) ||| Moldova   (Vasiliev & Mikhailov, 2023) ||| North Macedonia   (Komnenov, 2014) ||| Norway   (Aakra & Hauge, 2003) ||| Poland   (Rozwałka & Stanska, 2008) ||| Portugal   (Branco et al., 2019) ||| Romania   (no references) ||| Russia, Eastern   (no references) ||| Russia, Southern   (Otto, 2022; Ponomarev, 2022) ||| Serbia   (Grbić & Hänggi, 2023) ||| Slovenia   (Kostanjšek & Kuntner, 2015) ||| Spain   (Branco et al., 2019) ||| Sweden   (Kronestedt, 2001) ||| Switzerland   (no references) ||| Turkey (Asia)   (Danışman et al., 2024a) ||| Ukraine   (Polchaninova & Prokopenko, 2019) |||
References

Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129 pdf

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Bosmans R, Van Keer J, Russell-Smith A, Kronestedt T, Alderweireldt M, Bosselaers J, De Koninck H (2013) Spiders of Crete (Araneae). A catalogue of all currently known species from the Greek island of Crete. Nieuwsbrief van de Belgische Arachnologische Vereniging 28 (Suppl. 1): 1-147 pdf

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254 pdf

Danışman T, Kunt K B, Özkütük R S, Coşar İ (2024a) The checklist of the spiders of Turkey. Version 2024 [last updated 01 January 2024], online at http://www.spidersofturkey.info pdf

Grbac I, Katušić L, Lukić M (2019) Catalogue of spiders (Araneae) deposited in the Croatian Natural History Museum. Natura Croatica 28: 185-269 pdf

Grbić G, Hänggi A (2023) Additional spider records from Mt. Fruška Gora (Serbia) with a first insight into winter species. Acta Entomologica Serbica 27(2, 2022): 67-81 pdf

Hansen H (1995) Über die Arachniden-Fauna von urbanen Lebensräumen in Venedig - III. Die epigäischen Spinnen eines Stadtparkes (Arachnida: Araneae). Bollettino del Museo Civico di Storia Naturale di Venezia 44: 7-36 pdf

Komnenov M (2014) Spider fauna of the Osogovo Mt. Range, Northeastern Macedonia. Fauna Balkana 2: 1-267 pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Kronestedt T (1983) Spiders (Araneae) on the Great Alvar of the island of Öland, S Sweden. Entomologisk Tidskrift 104: 183-212 pdf

Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001] pdf

Logunov D V (1996b) A critical review of the spider genera Apollophanes O. P.-Cambridge, 1898 and Thanatus C. L. Koch, 1837 in North Asia (Araneae, Philodromidae). Revue Arachnologique 11: 133-202 pdf

Muster C, Thaler K (2003) The Thanatus striatus species group in the eastern Alps, with description of Thanatus firmetorum sp. n. (Araneae: Philodromidae). Bulletin of the British Arachnological Society 12: 376-382 pdf

Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.

Pantini P, Isaia M (2019) Araneae.it: the online catalog of Italian spiders, with addenda on other arachnid orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). Fragmenta Entomologica 51: 127-152 pdf

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Pozzi S, Hänggi A (1998) Araignées nouvelles ou peu connues de la Suisse (Arachnida: Araneae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 71: 33-47 pdf

Řezáč M, Růžička V (2022) Thanatus aridorum Šilhavý, 1940 from Czechia is a junior synonym of Thanatus formicinus (Clerck, 1757) (Araneae: Philodromidae). Arachnologische Mitteilungen 63: 15-18 pdf

Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Szita É, Samu F (2000) Taxonomical review of Thanatus species (Philodromidae, Araneae) of Hungary. Acta Zoologica Academiae Scientiarum Hungaricae 46: 155-179 pdf

Van Keer J, Van Keer K, De Koninck H, Ramel G (2010) The arachnofauna (Araneae) of wetland Kerkini (Macedonia-Northern Central Greece). Acta Zoologica Bulgarica 62: 141-160 pdf

Vasiliev A I, Mikhailov K G (2023) Annotated checklist of spiders (Araneae) of the Republic of Moldova. Invertebrate Zoology 20: 307-342 pdf

Vrenozi B, Jäger P (2012) A faunistic study on ground-dwelling spiders (Araneae) in the Tirana district, Albania. Arachnologische Mitteilungen 44: 81-87 pdf

WSC (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, online at http://wsc.nmbe.ch (22.01.2024) doi: 10.24436/2 pdf

Zarikian N (2021) A survey of running crab spiders Philodromidae (Araneae) of Armenia. Bulletin of the Iraq Natural History Museum 16: 495-508 pdf

Updates
05-02-2023 Distribution insert Detail
09-11-2022 Distribution update Detail
02-09-2022 Image insert
23-12-2021 Distribution insert Detail
26-07-2020 Image insert
24-09-2015 Distribution update Detail
12-11-2013 Image insert
15-10-2013 Distribution update Detail