Attulus penicillatus (Simon, 1875)
Description
Male
Palp male: dorsally with distinctly marked off colour pattern, tibia of palpal strongly thickened. Prosoma red-brown. Legs reddish yellow. Opisthosoma dark, provided with whitish grey hairs.
Body length male: 3.0-4.5 mmFemale
Copulatory openings of epigyne distinctly separated by narrow elevation at least over half of length of epigyne.
Body length female: 4.4-4.6 mmAdditional information
In warm places on sandy ground
Distribution
Figures
Pedipalp, ventral view
(Prószyński 1991)
Pedipalp, ventral view
(Żabka 1997)
Pedipalp
(Żabka 1997)
Pedipalp, ventral view
(Metzner 1999)
Pedipalp, retrolateral view
(Metzner 1999)
Pedipalp, ventral view
(Iorio 2023)
Habitus
(Żabka 1997)
Epigyne
(Żabka 1997)
Epigyne
(Metzner 1999)
Vulva
(Żabka 1997)
Vulva
(Metzner 1999)
Habitus
(Żabka 1997)
Distribution List
"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.
References
Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) 
Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510
Bosmans R, Chatzaki M (2005) A catalogue of the spiders of Greece – a critical review of all spider species cited from Greece with their localities. Nieuwsbrief van de Belgische arachnologische Vereniging 20 (2, suppl.): 1-124
Deltshev C D, Ćurčić B P M, Blagoev GA (2003b) The spiders of Serbia. Inst. Zoology, Fac. Biology, Belgrade & Inst. Zoology, Bulg. Acad. Sci., Sofia. 832 pp.
Esyunin S L, Efimik V E (1996a) Catalogue of the spiders (Arachnida, Aranei) of the Urals. KMK Scientific Press, Moscow, 228 pp.
Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp.
Helsdingen P J van, IJland S (2015) A quick scan of the spider fauna in the surroundings of Përmet, Albania (Arachnida, Araneae) - preliminary report. Nieuwsbrief SPINED 35: 15-33
Iorio E (2023) Les peuplements d’araignées (Araneae) de la réserve naturelle régionale des Isles du Drac (Isère, France), avec focus sur deux zones humides. Bulletin de l'Association Française d'Arachnologie 10: 4-40
Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204
Komnenov M (2014) Spider fauna of the Osogovo Mt. Range, Northeastern Macedonia. Fauna Balkana 2: 1-267
Maurer R, Hänggi A (1990) Katalog der schweizerischen Spinnen. Documenta Faunistica Helvetiae 12
Metzner H (1999) Die Springspinnen (Araneae, Salticidae) Griechenlands. Andrias 14: 1-279
Milosevic B (2002) Aranea – Popis vrsta – Checklist. Hrvatsko entomološko društvo – Inventar entomofaune Hrvatske [Aranea – List of species – Checklist. Croatian Entomological Society – Inventory of Croatian entomofauna].
Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.
Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App
Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp.
Prószyński J (1991) in Heimer S, Nentwig W: Spinnen Mitteleuropas. Parey Hamburg
Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html
Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html
Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184
Thaler K (1997c) Beiträge zur Spinnenfauna von Nordtirol - 4: Dionycha (Anyphaenidae, Clubionidae, Heteropodidae, Liocranidae, Philodromidae, Salticidae, Thomisidae, Zoridae). Veröffentlichungen des Museum Ferdinandeum in Innsbruck 77: 233-285
Weiss I, Petrișor A (1999) List of the spiders (Arachnida: Araneae) from Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 41: 79-107
WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (13.2.2026) doi: 10.24436/2
Żabka M (1997) Salticidae: Pajaki skaczace (Arachnida: Araneae) (Salticidae of Poland). Fauna Polski (Fauna Poloniae) 19: 1-188