Gnaphosidae: Micaria Westring, 1851
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1

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 Femur I dark brown, distal half light yellow with black-brown spot, a bristle within (often broken off); male pedipalp with narrow cymbium, ventrally with 2 and basally with one bristle; retinaculum narrow; epigyne with a bent ridge; receptaculum seminis oval, ducts divergent; pedipalp femur very large

Micaria dives (Lucas, 1846)
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male Pedipalp
(Thaler 1981c)

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male Pedipalp, ventral view
(Bosmans & Blick 2000)

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male Pedipalp
(Bosmans & Blick 2000)
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male Pedipalp
(Thaler 1981c)

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male Bulbus, ventral view
(Thaler 1981c)

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female Epigyne
(Bosmans & Blick 2000)
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female Epigyne
(Brændegaard 1966)

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female Epigyne
(Levy 1973)

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female Vulva, ventral view
(Thaler 1981c)
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female Vulva
(Bosmans & Blick 2000)

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female Vulva, ventral view
(Thaler 1981c)

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female Vulva
(Levy 1973)
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Habitus
(Brændegaard 1966)
- Femur I without black-brown spot on bright background; sexual organ different (cymbium only in subopaca with 2 distal bristles

  2  
2
(1)

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 Chelicerae with long grey hairs and with conspicuously, mostly dense copper-coloured scale hairs

Micaria fulgens (Walckenaer, 1802)
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male Pedipalp
(Reimoser 1937)

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male Pedipalp
(Roberts 1995)

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male Pedipalp
(Roberts 1995)
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male Cymbium
(Reimoser 1937)

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female Epigyne
(Brændegaard 1966)

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female Epigyne
(Miller 1971)
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female Vulva
(Miller 1971)

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female Vulva
(Reimoser 1937)
- Chelicerae only with grey, bristle-like hairs

  3  
3
(2)

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 Femur I and II distally with 2 bristles; male pedipalp with 1 or 2 small hump-shaped tibial apophysis; cymbium with 3 bristles; embolus very thick; retinaculum missing; epigyne without transverse ridge, medially with variable longitudinal bar

Micaria rossica Thorell, 1875
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male Pedipalp, ventral view
(Platnick & Shadab 1988)

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male Pedipalp
(Platnick & Shadab 1988)

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male Pedipalp
(Pesarini 2000)
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female Epigyne
(Platnick & Shadab 1988)

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female Epigyne
(Pesarini 2000)

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female Vulva
(Platnick & Shadab 1988)
- Femur I and II mostly with one dorsal bristle (only in aenea with 2 bristles); male pedipalp with 1 or 2 larger tibial apophyses; epigyne different

  4  
4
(3)

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 Tibial apophysis of male pedipalp dorsally bent; cymbium moderately long, distally with 4 bristles; embolus screw-like; retinaculum large; epigyne with glasses-like structure

Micaria guttulata (C. L. Koch, 1839)
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male Pedipalp, ventral view
(Miller 1967)

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male Pedipalp
(Miller 1967)

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male Pedipalp
(Roberts 1998)
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male Pedipalp
(Reimoser 1937)

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male Cymbium
(Reimoser 1937)

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female Epigyne
(Roberts 1998)
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female Epigyne
(Roberts 1998)

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female Vulva
(Miller 1971)

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female Vulva
(Reimoser 1937)
- Different

  5  
5
(4)

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 Femur I and II distally with 2 bristles; male pedipalp with reduced retinaculum; cymbium with 4 bristles; epigyne with large, anteriorly very broad groove

Micaria aenea Thorell, 1871
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male Pedipalp
(Miller 1967)

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male Pedipalp, ventral view
(Miller 1967)

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male Pedipalp, ventral view
(Platnick & Shadab 1988)
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male Pedipalp
(Platnick & Shadab 1988)

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female Epigyne
(Platnick & Shadab 1988)

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female Vulva
(Miller 1967)
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female Vulva
(Platnick & Shadab 1988)

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female Vulva, ventral view
(Tullgren 1946)
- Femur I and II with one dorsal bristle; retinaculum not reduced; epigyne different

  6  
6
(5)		 Cymbium distally with 3 bristles; epigyne with 2 shallow grooves, arcuately bordered or with one large groove and strongly sclerotised margin

  7  
- Cymbium distally with 4 bristles; epigyne without or with small groove

  9  
7
(6)

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 Femur III and IV with one distal bristle in basal half; male pedipalp as fig.; epigyne with large, deep groove, its anterior margin strongly sclerotised

Micaria alpina L. Koch, 1872
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male Pedipalp, ventral view
(Platnick & Shadab 1988)

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male Pedipalp, ventral view
(Locket & Millidge 1951)

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male Pedipalp
(Platnick & Shadab 1988)
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male Pedipalp
(Locket & Millidge 1951)

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female Epigyne
(Platnick & Shadab 1988)

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female Epigyne
(Locket & Millidge 1951)
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female Epigyne
(Locket et al. 1974)

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female Epigyne
(Locket et al. 1974)

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female Vulva
(Platnick & Shadab 1988)
- Femur III and IV with 2 distal bristles; sexual organ different

  8  
8
(7)

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 Male pedipalp with long and narrow tibial apophysis; embolus thick; distal retinaculum larger; epigyne with shallow groove and V-shaped structure (transverse ridges) in the centre

Micaria lenzi Bösenberg, 1899
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male Pedipalp
(Roberts 1998)

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male Pedipalp
(Reimoser 1937)

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male Cymbium
(Reimoser 1937)
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female Epigyne
(Roberts 1998)

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female Vulva
(Reimoser 1937)
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 Male pedipalp with shorter tibial apophysis; embolus thin; retinaculum small, in the middle of bulbus; epigyne as fig.

Micaria silesiaca L. Koch, 1875
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male Pedipalp
(Locket et al. 1974)

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male Pedipalp, ventral view
(Miller 1971)

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male Pedipalp, ventral view
(Locket et al. 1974)
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male Pedipalp
(Reimoser 1937)

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female Epigyne
(Miller 1971)

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female Epigyne
(Locket & Millidge 1953)
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female Vulva
(Thaler & Noflatscher 1990)

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female Vulva
(Miller 1967)

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female Vulva
(Reimoser 1937)
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female Vulva, dorsal view
(Tullgren 1946)
9
(6)		 Male pedipalp with 2 ± towering tibial apophysis; female tibia I and II ventrally with 1 or 2 bristles; sclerotised epigynal ridge very broad

  10  
- Male pedipalp with 1 tibial apophysis, appressed on cymbium; female tibia I and II ventrally without (pulicaria) or with 3 or 5 pairs of bristles respectively; sclerotised epigynal ridge narrow

  11  
10
(9)

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 Prosoma mostly distinctly red-brown (remove scale hairs), very elongated, 1.75-2.0 times as long as wide; opisthosoma ventrally without white longitudinal band, often with white dots; male pedipalp as fig.; position of cymbial bristles variable; epigyne with longish oval receptaculum seminis, reaching to transverse ridge at least

Micaria formicaria (Sundevall, 1831)
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male Pedipalp
(Azheganova 1968)

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male Pedipalp
(Pesarini unpubl.)

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male Pedipalp
(Roberts 1995)
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male Cymbium
(Reimoser 1937)

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female Epigyne
(Miller 1971)

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female Epigyne
(Pesarini unpubl.)
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female Vulva
(Azheganova 1968)

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female Vulva
(Miller 1967)

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female Vulva
(Reimoser 1937)
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female Vulva, dorsal view
(Tullgren 1946)

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female Vulva, ventral view
(Tullgren 1946)
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 Prosoma mostly black-brown (remove scale hairs), only 1.3-1.8 times as long as wide; opisthosoma ventrally with white longitudinal band (can be abraded); cymbial bristles variable; epigyne with egg-shaped receptaculum seminis, not reaching to transverse ridge

Micaria coarctata (Lucas, 1846)
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male Pedipalp, ventral view
(Bosmans & Blick 2000)

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male Pedipalp, ventral view
(Chyzer & Kulczyński 1897)

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male Pedipalp
(Levy 2002)
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male Pedipalp
(Bosmans & Blick 2000)

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male Pedipalp
(Chyzer & Kulczyński 1897)

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male Pedipalp
(Levy 2002)
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female Epigyne
(Bosmans & Blick 2000)

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female Epigyne
(Chyzer & Kulczyński 1897)

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female Epigyne
(Levy 2002)
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female Epigyne
(Simon 1932)

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female Vulva
(Bosmans & Blick 2000)

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female Vulva, dorsal view
(Tullgren 1946)
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female Spermatheca
(Levy 2002)

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Prosoma, dorsal view
(Levy 2002)

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Eyes, dorsal view
(Levy 2002)
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Opisthosoma, dorsal view
(Levy 2002)
11
(9)

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 Male pedipalp with very elongated retinaculum; pedipalp tibia dorsally sometimes with hump; female tibia I and II ventrally without bristles

Micaria pulicaria (Sundevall, 1831)
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male Pedipalp, ventral view
(Platnick & Shadab 1988)

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male Pedipalp, retrolateral view
(Platnick & Shadab 1988)

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female Epigyne, dorsal view
(Kovblyuk & Nadolny 2008)
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female Epigyne, ventral view
(Kovblyuk & Nadolny 2008)

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female Vulva
(Platnick & Shadab 1988)

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female Vulva, ventral view
(Tullgren 1946)
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Femur IV, dorsal view
(Muster & Michalik 2020)
- Male pedipalp with distinctly shorter retinaculum; female tibia I and II ventrally with 3 or 5-6 pairs of bristles

  12  
12
(11)

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 Male pedipalp with very strongly bent embolus, without excrescence between embolus and retinaculum; female tibia I and II ventrally with 5-6 pairs of thin bristles; epigyne variable, with bent, mostly bipartite transverse ridge, without groove, posteriorly with sickle-shaped structure

Micaria nivosa L. Koch, 1866
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male Pedipalp
(Mikhailov 1988)

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male Pedipalp, ventral view
(Aakra unpubl.)

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male Pedipalp, ventral view
(Miller 1967)
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male Pedipalp, ventral view
(Mikhailov 1988)

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male Pedipalp, lateral view
(Aakra unpubl.)

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male Pedipalp
(Miller 1967)
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male Pedipalp
(Mikhailov 1988)

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female Epigyne
(Miller 1971)

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female Vulva
(Miller 1967)
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female Vulva, dorsal view
(Tullgren 1946)
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 Embolus and retinaculum less bent, with a excrescence between embolus and retinaculum; female tibia I and II ventrally with 3 pairs of robust bristles; epigyne with a transverse ridge within a groove, posteriorly without sickle-shaped structure

Micaria albovittata (Lucas, 1846)
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male Pedipalp, ventral view
(Bosmans & Blick 2000)

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male Pedipalp, ventral view
(Levy 2002)

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male Pedipalp, ventral view
(Locket & Millidge 1951)
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male Pedipalp
(Bosmans & Blick 2000)

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male Pedipalp, retrolateral view
(Levy 2002)

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male Pedipalp
(Locket & Millidge 1951)
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male Pedipalp
(Roberts 1995)

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female Epigyne
(Bosmans & Blick 2000)

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female Epigyne
(Levy 2002)
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female Epigyne
(Roberts 1995)

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female Epigyne
(Locket & Millidge 1951)

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female Vulva
(Bosmans & Blick 2000)
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female Vulva
(Miller 1967)

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female Spermathecae, dorsal view
(Levy 2002)

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Prosoma, dorsal view
(Levy 2002)
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Eyes, dorsal view
(Levy 2002)

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Eyes, dorsal view
(Locket & Millidge 1951)

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Metatarsus and tarsus
(Locket & Millidge 1951)
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Opisthosoma, dorsal view
(Levy 2002)

Further taxa

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1. Micaria belezma Bosmans, 2000
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male Pedipalp, retrolateral view
(Bosmans & Blick 2000)

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male Pedipalp, ventral view
(Bosmans & Blick 2000)

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2. Micaria bosmansi Kovblyuk & Nadolny, 2008
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male Pedipalp, retrolateral view
(Kovblyuk & Nadolny 2008)

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male Pedipalp, ventral view
(Kovblyuk & Nadolny 2008)

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male Pedipalp, ventral view
(Ponomarev et al. 2018a)
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male Bulbus, ventral view
(Kovblyuk & Nadolny 2008)

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male Median apophysis
(Kovblyuk & Nadolny 2008)

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male Palpal tibia, retrolateral view
(Kovblyuk & Nadolny 2008)
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female Epigyne, ventral view
(Kovblyuk & Nadolny 2008)

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female Epigyne, ventral view
(Kovblyuk & Nadolny 2008)

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female Vulva, ventral view
(Kovblyuk & Nadolny 2008)
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female Vulva, dorsal view
(Kovblyuk & Nadolny 2008)

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3. Micaria brignolii (Bosmans & Blick, 2000)
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male Pedipalp, ventral view
(Bosmans & Blick 2000)

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male Pedipalp
(Bosmans & Blick 2000)

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male Pedipalp, retrolateral view
(Barrientos et al. 2022)
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female Epigyne
(Bosmans & Blick 2000)

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female Epigyne
(Barrientos et al. 2022)

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female Vulva
(Bosmans & Blick 2000)

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4. Micaria constricta Emerton, 1894
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male Pedipalp, ventral view
(Platnick & Shadab 1988)

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male Pedipalp, retrolateral view
(Platnick & Shadab 1988)

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male Pedipalp, retrolateral view
(Holm 1958b)
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male Pedipalp, ventral view
(Holm 1958b)

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male Pedipalp, ventral view
(Løvbrekke unpubl.)

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male Embolic division, retrolateral view
(Løvbrekke unpubl.)
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female Epigyne
(Platnick & Shadab 1988)

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female Epigyne, ventral view
(Holm 1958b)

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female Epigyne
(Løvbrekke unpubl.)
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female Vulva, dorsal view
(Platnick & Shadab 1988)

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female Vulva
(Holm 1958b)

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female Vulva, dorsal view
(Løvbrekke unpubl.)

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5. Micaria corvina Simon, 1878
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male Pedipalp, prolateral view
(Levy 2002)

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male Pedipalp, retrolateral view
(Levy 2002)

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male Pedipalp, ventral view
(Levy 2002)
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female Epigyne
(Levy 2002)

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female Vulva, dorsal view
(Levy 2002)

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6. Micaria cyrnea Brignoli, 1983
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male Pedipalp, ventral view
(Bosmans & Blick 2000)

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male Pedipalp
(Bosmans & Blick 2000)

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female Epigyne
(Bosmans & Blick 2000)
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female Epigyne
(Bosmans & Blick 2000)

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female Vulva, ventral view
(Hansen 1992)

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7. Micaria donensis Ponomarev & Tsvetkov, 2006
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male Pedipalp, ventral view
(Ponomarev & Tsvetkov 2006)

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male Pedipalp, lateral view
(Ponomarev & Tsvetkov 2006)

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female Epigyne, ventral view
(Ponomarev & Tsvetkov 2006)

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8. Micaria funerea Simon, 1878
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male Pedipalp
(Simon 1932)

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female Epigyne
(Simon 1932)

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9. Micaria gulliae Tuneva & Esyunin, 2003
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male Pedipalp, dorsal view
(Tuneva & Esyunin 2003a)

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male Pedipalp, ventral view
(Tuneva & Esyunin 2003a)

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male Pedipalp, lateral view
(Tuneva & Esyunin 2003a)
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female Epigyne
(Tuneva & Esyunin 2003a)

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female Vulva
(Tuneva & Esyunin 2003a)

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10. Micaria guttigera Simon, 1878
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male Pedipalp
(Simon 1932)

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female Epigyne
(Simon 1932)

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11. Micaria ignea O. Pickard-Cambridge, 1872
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male Pedipalp, ventral view
(Chatzaki et al. 2002a)

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male Pedipalp, retrolateral view
(Chatzaki et al. 2002a)

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male Pedipalp, ventral view
(Bosmans & Blick 2000)
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male Pedipalp, retrolateral view
(Bosmans & Blick 2000)

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female Epigyne, ventral view
(Chatzaki et al. 2002a)

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female Epigyne, ventral view
(Bosmans & Blick 2000)
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female Vulva, dorsal view
(Chatzaki et al. 2002a)

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female Vulva
(Bosmans & Blick 2000)

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12. Micaria kopetdaghensis Mikhailov, 1986
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male Pedipalp, ventral view
(Mikhailov 1988)

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male Pedipalp, ventral view
(Mikhailov 1988)

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male Pedipalp, retrolateral view
(Mikhailov 1988)
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male Pedipalp, prolateral view
(Mikhailov 1988)

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male Palpal tibia, dorsal view
(Mikhailov 1988)

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female Epigyne, ventral view
(Mikhailov 1988)
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female Epigyne, ventral view
(Mikhailov 1988)

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13. Micaria marchesii (Caporiacco, 1936)
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female Epigyne
(Caporiacco 1936a)

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14. Micaria micans (Blackwall, 1858)
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male Pedipalp
(Brændegaard 1966)

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male Pedipalp, ventral view
(Brændegaard 1966)

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male Pedipalp, ventral view
(Locket & Millidge 1951)
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male Pedipalp
(Locket & Millidge 1951)

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male Pedipalp, retrolateral view
(Kovblyuk & Nadolny 2008)

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male Pedipalp, ventral view
(Kovblyuk & Nadolny 2008)
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male Pedipalp, ventral view
(Heimer & Nentwig 1991)

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male Pedipalp, ventral view
(Heimer & Nentwig 1991)

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male Pedipalp, retrolateral view
(Heimer & Nentwig 1991)
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male Pedipalp, prolateral view
(Muster & Michalik 2020)

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male Pedipalp, retrolateral view
(Muster & Michalik 2020)

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male Pedipalp, ventral view
(Muster & Michalik 2020)
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male Pedipalp, prolateral view
(Dolejš 2020)

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male Pedipalp, ventral view
(Tuneva 2007)

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male Bulbus, ventral view
(Kovblyuk & Nadolny 2008)
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male Median apophysis
(Kovblyuk & Nadolny 2008)

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male Palpal tibia, retrolateral view
(Kovblyuk & Nadolny 2008)

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female Epigyne
(Brændegaard 1966)
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female Epigyne
(Locket & Millidge 1951)

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female Epigyne
(Heimer & Nentwig 1991)

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female Epigyne
(Heimer & Nentwig 1991)
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female Epigyne
(Tuneva 2007)

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female Epigyne
(Langraf et al. 2024)

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female Vulva
(Miller 1967)
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female Vulva, ventral view
(Tullgren 1946)

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female Vulva, schematically
(Muster & Michalik 2020)

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female Vulva
(Muster & Michalik 2020)
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female Vulva
(Muster & Michalik 2020)

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female Vulva, dorsal view
(Tuneva 2007)

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Eyes, dorsal view
(Locket & Millidge 1951)
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Femur IV, dorsal view
(Muster & Michalik 2020)

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15. Micaria pallipes (Lucas, 1846)
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male Pedipalp, ventral view
(Bosmans 1999)

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male Pedipalp, ventral view
(Levy 2002)

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male Pedipalp, ventral view
(Mikhailov 1988)
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male Pedipalp
(Bosmans & Blick 2000)

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male Pedipalp, retrolateral view
(Levy 2002)

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male Pedipalp
(Mikhailov 1988)
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male Pedipalp, dorsal view
(Tuneva 2007)

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male Pedipalp, retrolateral view
(Tuneva 2007)

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male Pedipalp, ventral view
(Tuneva 2007)
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male Tibial apophysis
(Mikhailov 1988)

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female Epigyne
(Bosmans & Blick 2000)

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female Epigyne
(Levy 2002)
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female Epigyne
(Tuneva 2007)

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female Vulva
(Bosmans & Blick 2000)

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female Vulva, dorsal view
(Tuneva 2007)
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female Spermathecae, dorsal view
(Levy 2002)

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Opisthosoma, dorsal view
(Levy 2002)

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Habitus, dorsal view
(Tuneva 2007)

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16. Micaria palmgreni Wunderlich, 1980
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female Pedipalp, retrolateral view
(Wunderlich 1980h)

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female Epigyne, ventral view
(Wunderlich 1980h)

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female Vulva, dorsal view
(Wunderlich 1980h)

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17. Micaria sociabilis Kulczyński, 1897
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male Pedipalp, ventral view
(Bosmans & Blick 2000)

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male Pedipalp
(Bosmans & Blick 2000)

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male Pedipalp
(Simon 1932)
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male Pedipalp, retrolateral view
(Pfliegler 2014)

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male Pedipalp, ventral view
(Pfliegler 2014)

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male Pedipalp, prolateral view
(Pfliegler 2014)
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male Pedipalp, ventral view
(Mikhailov 2016)

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male Pedipalp, retrolateral view
(Mikhailov 2016)

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male Pedipalp, ventral view
(Ponomarev et al. 2016)
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male Tibial apophysis
(Heimer & Nentwig 1991)

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male Tibial apophysis
(Heimer & Nentwig 1991)

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male Tibial apophysis
(Mikhailov 2016)
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male Tibial apophysis
(Mikhailov 2016)

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female Epigyne
(Bosmans & Blick 2000)

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female Epigyne
(Heimer & Nentwig 1991)
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female Epigyne/vulva
(Pfliegler 2014)

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female Epigyne
(Ponomarev et al. 2016)

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female Vulva
(Bosmans & Blick 2000)

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18. Micaria subopaca Westring, 1861
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male Pedipalp
(Brændegaard 1966)

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male Pedipalp
(Miller 1967)

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male Pedipalp, ventral view
(Brændegaard 1966)
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male Pedipalp, ventral view
(Miller 1967)

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male Pedipalp, ventral view
(Locket & Millidge 1953)

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male Pedipalp
(Roberts 1995)
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male Pedipalp
(Locket & Millidge 1953)

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male Pedipalp, ventral view
(Heimer & Nentwig 1991)

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male Pedipalp, retrolateral view
(Heimer & Nentwig 1991)
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male Pedipalp, ventral view
(Løvbrekke unpubl.)

 speciesimage

male Pedipalp, prolateral view
(Tuneva 2007)

 speciesimage

male Pedipalp, retrolateral view
(Tuneva 2007)
speciesimage

male Pedipalp, ventral view
(Tuneva 2007)

 speciesimage

male Pedipalp, retrolateral view
(Seropian et al. 2023b)

 speciesimage

male Pedipalp, ventral view
(Seropian et al. 2023b)
speciesimage

male Cymbium
(Reimoser 1937)

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male Palpal tibia, retrolateral view
(Løvbrekke unpubl.)

 speciesimage

female Epigyne
(Brændegaard 1966)
speciesimage

female Epigyne
(Miller 1967)

 speciesimage

female Epigyne
(Locket & Millidge 1953)

 speciesimage

female Epigyne
(Heimer & Nentwig 1991)
speciesimage

female Epigyne/vulva
(Pfliegler 2014)

 speciesimage

female Epigyne
(Ponomarev et al. 2016)

 speciesimage

female Epigyne
(Løvbrekke unpubl.)
speciesimage

female Epigyne
(Tuneva 2007)

 speciesimage

female Vulva
(Miller 1967)

 speciesimage

female Vulva
(Reimoser 1937)
speciesimage

female Vulva
(Tullgren 1946)

 speciesimage

female Vulva, dorsal view
(Løvbrekke unpubl.)

 speciesimage

female Vulva, dorsal view
(Tuneva 2007)
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Leg II, lateral view
(Tuneva 2007)

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19. Micaria triguttata Simon, 1884
speciesimage

male Pedipalp, retrolateral view
(Wunderlich 1980h)

 speciesimage

male Pedipalp, ventral view
(Wunderlich 1980h)

 speciesimage

male Pedipalp, prolateral view
(Wunderlich 1980h)
speciesimage

male Palpal tibia, dorsal view
(Wunderlich 1980h)

 speciesimage

female Epigyne, ventral view
(Bosmans & Blick 2000)

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female Opisthosoma, dorsal view
(Ledoux et al. 2003)

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cp

20. Micaria tripunctata Holm, 1978
speciesimage

male Pedipalp
(Platnick & Shadab 1988)

 speciesimage

male Pedipalp
(Platnick & Shadab 1988)

 speciesimage

male Pedipalp, retrolateral view
(Pantini et al. 2020b)
speciesimage

male Pedipalp, ventral view
(Pantini et al. 2020b)

 speciesimage

male Pedipalp, ventral view
(Løvbrekke unpubl.)

 speciesimage

male Pedipalp, retrolateral view
(Tuneva 2007)
speciesimage

male Pedipalp, ventral view
(Tuneva 2007)

 speciesimage

female Epigyne
(Platnick & Shadab 1988)

 speciesimage

female Epigyne
(Pantini et al. 2020b)
speciesimage

female Epigyne
(Løvbrekke unpubl.)

 speciesimage

female Epigyne
(Tuneva 2007)

 speciesimage

female Vulva
(Platnick & Shadab 1988)
speciesimage

female Vulva, dorsal view
(Pantini et al. 2020b)

 speciesimage

female Vulva, dorsal view
(Løvbrekke unpubl.)

 speciesimage

female Vulva, dorsal view
(Tuneva 2007)
speciesimage

Habitus, dorsal view
(Tuneva 2007)

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21. Micaria zonsteini (Mikhailov, 2016)
speciesimage

male Pedipalp, ventral view
(Mikhailov 2016)

 speciesimage

male Pedipalp, retrolateral view
(Mikhailov 2016)

 speciesimage

male Pedipalp, prolateral view
(Mikhailov 2016)
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male Tibial apophysis, retrolateral view
(Mikhailov 2016)

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female Epigyne
(Mikhailov 2016)

 speciesimage

female Vulva
(Mikhailov 2016)
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female Sternum
(Mikhailov 2016)