Salticidae  Blackwall, 1841 / Attulus  Simon, 1889   23 species

Attulus penicillatus (Simon, 1875)

Description
Male

Palp male: dorsally with distinctly marked off colour pattern, tibia of palpal strongly thickened. Prosoma red-brown. Legs reddish yellow. Opisthosoma dark, provided with whitish grey hairs.

Body length male: 3.0-4.5 mm
Female

Copulatory openings of epigyne distinctly separated by narrow elevation at least over half of length of epigyne. 

Body length female: 4.4-4.6 mm
Additional information

In warm places on sandy ground

Distribution
  Presence
  No data
Global distribution (WSC 2026): Europe, Caucasus, Russia (Europe to Far East), Kazakhstan, China, Korea, Japan
Figures
Distribution List
Albania   (van Helsdingen & IJland, 2015) ||| Austria   (Thaler, 1997c) ||| Azerbaijan   (Otto, 2022) ||| Bulgaria   (Blagoev et al., 2018) ||| Croatia   (Milosevic, 2002) ||| Czechia   (Růžička & Řezáč, 2022a) ||| France   (Iorio, 2023) ||| Georgia   (Otto, 2022) ||| Germany   (Blick et al., 2016) ||| Greece   (Bosmans & Chatzaki, 2005) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Isaia et al., 2025) ||| North Macedonia   (Komnenov, 2014) ||| Poland   (Rozwałka & Stanska, 2008) ||| Romania   (Weiss & Petrișor, 1999) ||| Russia, Central   (Ponomarev, 2022) ||| Russia, Eastern   (Esyunin & Efimik, 1996a) ||| Russia, Southern   (Otto, 2022; Ponomarev, 2022) ||| Serbia   (Deltshev et al., 2003b) ||| Slovakia   (Gajdoš et al., 1999) ||| Switzerland   (Maurer & Hänggi, 1990) ||| Ukraine   (Polchaninova & Prokopenko, 2019) |||
References

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510 pdf

Bosmans R, Chatzaki M (2005) A catalogue of the spiders of Greece – a critical review of all spider species cited from Greece with their localities. Nieuwsbrief van de Belgische arachnologische Vereniging 20 (2, suppl.): 1-124 pdf

Deltshev C D, Ćurčić B P M, Blagoev GA (2003b) The spiders of Serbia. Inst. Zoology, Fac. Biology, Belgrade & Inst. Zoology, Bulg. Acad. Sci., Sofia. 832 pp.

Esyunin S L, Efimik V E (1996a) Catalogue of the spiders (Arachnida, Aranei) of the Urals. KMK Scientific Press, Moscow, 228 pp. pdf

Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp.

Helsdingen P J van, IJland S (2015) A quick scan of the spider fauna in the surroundings of Përmet, Albania (Arachnida, Araneae) - preliminary report. Nieuwsbrief SPINED 35: 15-33 pdf

Iorio E (2023) Les peuplements d’araignées (Araneae) de la réserve naturelle régionale des Isles du Drac (Isère, France), avec focus sur deux zones humides. Bulletin de l'Association Française d'Arachnologie 10: 4-40 pdf

Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204 pdf

Komnenov M (2014) Spider fauna of the Osogovo Mt. Range, Northeastern Macedonia. Fauna Balkana 2: 1-267 pdf

Maurer R, Hänggi A (1990) Katalog der schweizerischen Spinnen. Documenta Faunistica Helvetiae 12 pdf

Metzner H (1999) Die Springspinnen (Araneae, Salticidae) Griechenlands. Andrias 14: 1-279 pdf

Milosevic B (2002) Aranea – Popis vrsta – Checklist. Hrvatsko entomološko društvo – Inventar entomofaune Hrvatske [Aranea – List of species – Checklist. Croatian Entomological Society – Inventory of Croatian entomofauna]. pdf

Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Prószyński J (1991) in Heimer S, Nentwig W: Spinnen Mitteleuropas. Parey Hamburg pdf

Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Thaler K (1997c) Beiträge zur Spinnenfauna von Nordtirol - 4: Dionycha (Anyphaenidae, Clubionidae, Heteropodidae, Liocranidae, Philodromidae, Salticidae, Thomisidae, Zoridae). Veröffentlichungen des Museum Ferdinandeum in Innsbruck 77: 233-285 pdf

Weiss I, Petrișor A (1999) List of the spiders (Arachnida: Araneae) from Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 41: 79-107 pdf

WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (20.5.2026) doi: 10.24436/2 pdf

Żabka M (1997) Salticidae: Pajaki skaczace (Arachnida: Araneae) (Salticidae of Poland). Fauna Polski (Fauna Poloniae) 19: 1-188 pdf

Updates
12-11-2025 Distribution update Detail
08-05-2023 Image insert
08-05-2023 Distribution update Detail
12-01-2023 Distribution update Detail
03-05-2018 Image insert
24-12-2015 Distribution update Detail