Thomisidae  Sundevall, 1833 / Xysticus  C. L. Koch, 1835   82 species

Xysticus luctator L. Koch, 1870

Description
Male

Male palp dark brown. Median bulbal apophysis with 2 tips arranged rectangularly. Tibial apophysis bipartite, massive ventral and inconspicuous retrolateral apophysis. Male: black-brown, eye field bright, 2 dark brown moon-like spots (sometimes 2 additional bright ones) on posterior margin of prosoma. Sternum longish oval to shield-shaped, brown to light brown with cream spots. Chelicerae brown with bright transverse band and one longer spine (female frontally many short spines). Legs dark brown, contrast-rich with white spots on junction femur/patella, distal leg segments brighter. Females overall with inconspicuous, bright legs. Metatarsus I and II and tibia with strikingly high number of spines. Opisthosoma female: beige, without pattern. Males: brown to dark brown, with thin cream margin and white, interrupted transverse bands in posterior third.

Body length male: 5.9-8.4 mm
Female

Epigyne with wide-oval epigynal groove, with broad, protruding septum. Vulva with anterior part consisting of 2 funnel-shaped, very dark parts, broadened posteriorly. Prosoma strongly bulged, round. Female: light beige to sandy-brown without distinct pattern, but with 2 dark moon-like spots on posterior margin of prosoma. 

Body length female: 7.0-13.0 mm
Additional information

from lowland to mountainous level, in habitats with poor vegetation

Distribution
  Presence
  No data
Global distribution (WSC 2026): Europe, Caucasus, Russia (Europe to South Siberia), Kazakhstan
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List

"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.

Austria   (Thaler & Knoflach, 2004b) ||| Azerbaijan   (Otto, 2022) ||| Belarus   (Ivanov, 2013) ||| Belgium   (Bosmans & Van Keer, 2017) ||| Bosnia and Herzegovina   (Komnenov, 2010) ||| Bulgaria   (Blagoev et al., 2018) ||| Czechia   (Růžička & Řezáč, 2022a) ||| Denmark   (Scharff & Gudik-Sørensen, 2006) ||| Estonia   (Vilbaste, 1987) ||| Finland   (Koponen et al., 2016) ||| France   (Le Peru, 2007) ||| Georgia   (Otto, 2022) ||| Germany   (Blick et al., 2016) ||| Greece   (Van Keer et al., 2010) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Isaia et al., 2025) ||| Luxembourg   (Kreuels et al., 2019) ||| Moldova   (Vasiliev & Mikhailov, 2023) ||| Netherlands   (van Helsdingen, 2025a) ||| North Macedonia   (Komnenov, 2014) ||| Norway   (Aakra et al., 2016) ||| Poland   (Rozwałka & Stanska, 2008) ||| Romania   (Weiss & Petrișor, 1999) ||| Russia, Central   (Ponomarev, 2022) ||| Russia, Eastern   (Esyunin & Efimik, 1996a) ||| Russia, Southern   (Otto, 2022; Ponomarev, 2022) ||| Russia, Western   (no references) ||| Serbia   (Grbić et al., 2021) ||| Slovakia   (Gajdoš et al., 1999) ||| Sweden   (Kronestedt, 2001) ||| Switzerland   (Maurer & Hänggi, 1990) ||| Ukraine   (Polchaninova & Prokopenko, 2019) ||| United Kingdom   (Lavery, 2019) |||
References

Aakra K, Morka G H, Antonson A, Farlund M, Wrånes R E, Frølandshagen R, Løvbrekke H, Furuseth P, Fjellberg A, Lemke M, Pfliegler W P, Andersen S, Olsen K M, Aadland B, Berggren K (2016) Spiders new to Norway (Arachnida, Araneae) with ecological, taxonomical and faunistic comments. Norwegian Journal of Entomology 63: 6-43 pdf

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510 pdf

Bosmans R, Van Keer K (2017) Een herziene soortenlijst van de Belgische spinnen (Araneae). Nieuwsbrief van de Belgische Arachnologische Vereniging 32: 39-69 pdf

Brændegaard J (1966) Edderkopper: Eller Spindlere I Danmarks Fauna 72: 1-224 pdf

Esyunin S L, Efimik V E (1996a) Catalogue of the spiders (Arachnida, Aranei) of the Urals. KMK Scientific Press, Moscow, 228 pp. pdf

Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp.

Grbić G, Hänggi A, Krnjajić S (2021) Spiders (Araneae) of Subotica Sandland (Serbia): additional arguments in environmental protection. Acta Zoologica Academiae Scientiarum Hungaricae 67: 15-61 pdf

Heimer S, Nentwig W (1991) Spinnen Mitteleuropas. Paul Parey Berlin pdf

Helsdingen P J van (2025a) Catalogus van de Nederlandse spinnen. Versie 2025.1. 249 pp. pdf

Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204 pdf

Ivanov V V (2013) The checklist of Belarusian spiders (Arachnida, Araneae). Zoology and Ecology 4: 293-311 pdf

Jantscher E (2001a) Revision der Krabbenspinnengattung Xysticus C.L. Koch, 1835 (Araneae, Thomisidae) in Zentraleuropa. Dissertation, Univ. Graz, 328 pp., 81 pls. pdf

Komnenov M (2010) Checklist of spiders (Araneae) of Bosnia and Herzegovina. Uzizaž i Biospeld 5(2009): 52-69 pdf

Komnenov M (2014) Spider fauna of the Osogovo Mt. Range, Northeastern Macedonia. Fauna Balkana 2: 1-267 pdf

Koponen S, Fritzén N R, Pajunen T (2016) Checklist of spiders in Finland (Araneae). 6th version, December 2016, online at http://biolcoll.utu.fi/arach/checklist_of_spiders_in_Finland.htm pdf

Kreuels M, Staudt A, Christian S (2019) Die Spinnenfauna von Luxemburg – eine Zusammenstellung der Nachweise aus den Jahren 1906-2018 (Arachnida: Araneae). Bulletin de la Société des naturalistes luxembourgeois 121: 203-230 pdf

Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001] pdf

Lavery A (2019) A revised checklist of the spiders of Great Britain and Ireland. Arachnology 18: 196-212 pdf

Le Peru B (2007) Catalogue et répartition des araignées de France. Revue Arachnologique 16: 1-468 pdf

Locket G H, Millidge A F (1951) British spiders. Vol. I. Ray Society, London, 310 pp. pdf

Locket G H, Millidge A F, Merrett P (1974) British Spiders, Volume III. Ray Society, London, 315 pp. pdf

Løvbrekke H (unpubl.)

Maurer R, Hänggi A (1990) Katalog der schweizerischen Spinnen. Documenta Faunistica Helvetiae 12 pdf

Morka G H (unpubl.)

Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Scharff N, Gudik-Sørensen O (2006) Katalog over Danmarks edderkopper (Araneae). Catalogue of the spiders of Denmark (Araneae). Entomologiske Meddelelser 74: 3-71 pdf

Thaler K, Knoflach B (2004b) Zur Faunistik der Spinnen (Araneae) von Österreich: Gnaphosidae, Thomisidae (Dionycha pro parte). Linzer Biologische Beiträge 36: 417-484 pdf

Van Keer J, Van Keer K, De Koninck H, Ramel G (2010) The arachnofauna (Araneae) of wetland Kerkini (Macedonia-Northern Central Greece). Acta Zoologica Bulgarica 62: 141-160 pdf

Vasiliev A I, Mikhailov K G (2023) Annotated checklist of spiders (Araneae) of the Republic of Moldova. Invertebrate Zoology 20: 307-342 pdf

Vilbaste A (1987) Eesti Ämblikud (Aranei). Annoteeritud nimestik. Estonian spiders (Aranei). An annotated check list. Valgus, Tallinn. 113 pp. & 512 maps on 67 unpag. pages pdf

Vogels J, Hiddes C (2023) Xysticus luctator L. Koch, 1870 (Araneae, Thomisidae) na 131 jaar weer in Nederland aangetroffen [Xysticus luctator L. Koch, 1870 (Araneae, Thomisidae) found again in the Netherlands after 131 years]. Nieuwsbrief SPINED 40: 2-7 pdf

Weiss I, Petrișor A (1999) List of the spiders (Arachnida: Araneae) from Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 41: 79-107 pdf

WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (13.2.2026) doi: 10.24436/2 pdf

Updates
12-11-2025 Distribution update Detail
13-03-2023 Image insert
18-01-2023 Distribution update Detail
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11-08-2020 Distribution update Detail
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01-12-2013 Distribution update Detail