Haplodrassus umbratilis (L. Koch, 1866)
Description
Male palp with plate-like bulge of distal bulbal apophysis, situated basally. Tibial apophysis somewhat widened distally. Epigyne with slantingly forward-directed protrusion. Prosoma light brown. Chelicerae darker than prosoma. Distal leg segments darker than prosoma. Opisthosoma light brown.
Body length male: 3.9-6.0 mmBody length female: 5.1-7.5 mm
Additional information
in dry and open pine forests and heathland, on the ground and under stones during the day
Distribution
Phenology
| Jan | Feb | Mar | Apr | May | Jun | Jul | Aug | Sep | Oct | Nov | Dec |
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Figures
Pedipalp, ventral view
(Miller 1971)
Pedipalp, ventral view
(Locket et al. 1974)
Pedipalp
(Roberts 1995)
Pedipalp, ventral view
(Kovblyuk et al. 2012b)
Pedipalp, ventral view
(Ponomarev & Shmatko 2017)
Pedipalp, retrolateral view
(Fjellberg unpubl.)
Pedipalp, ventral view
(Fjellberg unpubl.)
Pedipalp, ventral view
(Løvbrekke unpubl.)
Bulbus, apical view
(Kovblyuk et al. 2012b)
Bulbus, ventral view
(Ponomarev & Shmatko 2017)
Bulbus, ventro-lateral view
(Ponomarev & Shmatko 2017)
Embolus
(Løvbrekke unpubl.)
Palpal tibia
(Locket et al. 1974)
Palpal tibia, dorsal view
(Løvbrekke unpubl.)
Tibial apophysis
(Miller & Buchar 1977)
Tibial apophysis, retrolateral view
(Kovblyuk et al. 2012b)
Tibial apophysis, dorsal view
(Ponomarev & Shmatko 2017)
Epigyne
(Brændegaard 1966)
Epigyne
(Roberts 1995)
Epigyne
(Locket et al. 1974)
Epigyne, ventral view
(Kovblyuk et al. 2012b)
Epigyne
(Ponomarev & Shmatko 2017)
Epigyne
(Ponomarev & Shmatko 2017)
Epigyne
(Løvbrekke unpubl.)
Vulva
(Miller & Buchar 1977)
Vulva, ventral view
(Tullgren 1946)
Vulva, dorsal view
(Kovblyuk et al. 2012b)
Vulva
(Ponomarev & Shmatko 2017)
Vulva
(Ponomarev & Shmatko 2017)
Vulva, dorsal view
(Løvbrekke unpubl.)
Habitus, dorsal view
(Fjellberg unpubl.)
Distribution List
"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.
References
Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129 
Arnold K (2001) Beitrag zur Spinnenfauna (Arachnida, Araneae) des Fürstentums Liechtenstein. Berichte der Botanisch-Zoologischen Gesellschaft Liechtenstein-Sargans-Werdenberg 28: 211-244
Blagoev G (2002) Check list of Macedonian spiders (Araneae). Acta Zoologica Bulgarica 54: 9-34
Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018)
Bosmans R, Van Keer K (2017) Een herziene soortenlijst van de Belgische spinnen (Araneae). Nieuwsbrief van de Belgische Arachnologische Vereniging 32: 39-69
Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254
Brændegaard J (1966) Edderkopper: Eller Spindlere I Danmarks Fauna 72: 1-224
Cera I (2018) The checklist of Latvian spiders (Arachnida: Araneae). Environmental and Experimental Biology 16: 139-152
Danışman T, Kunt K B, Özkütük R S, Coşar İ (2024) The checklist of the spiders of Turkey. Version 2024 [last updated 01 January 2024], online at http://www.spidersofturkey.info
Fjellberg A (unpubl.)
Hermann E (1998) Die Spinnen (Araneae) ausgewählter Halbtrockenrasen im Osten Luxemburgs. Bulletin de la Société des naturalistes Luxembourgeois 99: 189-199
Ivanov V V (2013) The checklist of Belarusian spiders (Arachnida, Araneae). Zoology and Ecology 4: 293-311
Koponen S, Fritzén N R, Pajunen T (2016) Checklist of spiders in Finland (Araneae). 6th version, December 2016, online at http://biolcoll.utu.fi/arach/checklist_of_spiders_in_Finland.htm
Kovblyuk M M, Kastrygina Z A, Omelko M M (2012b) A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae, Gnaphosidae). Zookeys 205: 59-89
Kreuels M, Staudt A, Christian S (2019) Die Spinnenfauna von Luxemburg – eine Zusammenstellung der Nachweise aus den Jahren 1906-2018 (Arachnida: Araneae). Bulletin de la Société des naturalistes luxembourgeois 121: 203-230
Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001]
Lavery A (2019) A revised checklist of the spiders of Great Britain and Ireland. Arachnology 18: 196-212
Locket G H, Millidge A F, Merrett P (1974) British Spiders, Volume III. Ray Society, London, 315 pp.
Løvbrekke H (unpubl.)
Miller F (1971) Pavouci-Araneida. Klíč zvířeny ČSSR 4: 51-306
Miller F, Buchar J (1977) Neue Spinnenarten aus den Gattungen Zelotes Distel und Haplodrassus Chamberlin (Araneae, Gnaphosidae). Acta Universitatis Carolinae Biologica 1974: 157-171
Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.
Pantini P, Isaia M (2019) Araneae.it: the online catalog of Italian spiders, with addenda on other arachnid orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). Fragmenta Entomologica 51: 127-152
Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App
Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp.
Ponomarev A V, Shmatko V Y (2017) A new species of spiders of the genus Haplodrassus Chamberlin, 1922 (Aranei: Gnaphosidae) from the northern pre-Caspian area. Caucasian Entomological Bulletin 13: 11-14
Roberts M J (1995) Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp.
Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html
Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html
Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184
Tullgren A (1946) Svensk spindelfauna. 3. Egentiliga spinddlar. Araneae Fam 5-7. Clubionidae, Zoridae och Gnaphosidae. Stockholm (Entomol Fören.): 141 pp.
Ukhova N, Esyunin S, Sozontov A (2024) Spiders (Arachnida, Araneae) of the Visimskiy Biosphere Reserve (Middle Urals): 37 years of arachnological research. Biodiversity Data Journal 12(e114930): 1-37
Vasiliev A I, Mikhailov K G (2023) Annotated checklist of spiders (Araneae) of the Republic of Moldova. Invertebrate Zoology 20: 307-342
Vilbaste A (1987) Eesti Ämblikud (Aranei). Annoteeritud nimestik. Estonian spiders (Aranei). An annotated check list. Valgus, Tallinn. 113 pp. & 512 maps on 67 unpag. pages
WSC (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, online at http://wsc.nmbe.ch (22.01.2024) doi: 10.24436/2