Gnaphosidae  Banks, 1892 / Scotophaeus  Simon, 1893   21 species

Scotophaeus scutulatus (L. Koch, 1866)

Description

Tibial apophysis with distinct tip. Epigyne with ± square groove, distinctly longer than wide. Prosoma light to dark red-brown. Opisthosoma light to dark red-brown.

Body length male: 6.8-10.9 mm
Body length female: 8.3-15.9 mm
Additional information

mostly in and on buildings, rare in natural environment

Distribution
  Presence
  No data
Global distribution (WSC 2026): Europe, Algeria, Turkey, Caucasus, Russia (Europe to South Siberia), Iran, Central Asia
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List
Albania   (Deltshev et al., 2011b; Vrenozi, 2012; Blick, 2018) ||| Algeria   (Benhacene et al., 2023) ||| Austria   (Thaler & Knoflach, 2004b) ||| Azerbaijan   (Otto, 2022) ||| Belgium   (Bosmans & Van Keer, 2017) ||| Bosnia and Herzegovina   (Komnenov, 2010) ||| Bulgaria   (Blagoev et al., 2018) ||| Croatia   (Milosevic, 2002) ||| Cyprus   (Bosmans et al., 2019c) ||| Czechia   (Růžička & Řezáč, 2022a) ||| Denmark   (Scharff & Gudik-Sørensen, 2006) ||| France   (Le Peru, 2007) ||| Georgia   (Otto, 2022) ||| Germany   (Blick et al., 2016) ||| Greece   (Bosmans & Chatzaki, 2005) ||| Greece / Crete   (Bosmans & Chatzaki, 2005) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Isaia et al., 2025) ||| Italy / Sardinia   (Isaia et al., 2025) ||| Italy / Sicily   (Isaia et al., 2025) ||| Liechtenstein   (Arnold, 2001) ||| Lithuania   (Biteniekytė & Rėlys, 2012) ||| Luxembourg   (Christian, 2015; Kreuels et al., 2019) ||| Malta   (Pfliegler et al., 2017) ||| Moldova   (Vasiliev & Mikhailov, 2023) ||| Netherlands   (van Helsdingen, 2025a) ||| North Macedonia   (Deltshev et al., 2013) ||| Poland   (Rozwałka & Stanska, 2008) ||| Portugal   (Branco et al., 2019) ||| Romania   (Weiss & Petrișor, 1999) ||| Russia, Central   (no references) ||| Russia, Southern   (Otto, 2022; Ponomarev, 2022) ||| Serbia   (Stanković, 2023) ||| Slovakia   (Gajdoš et al., 1999) ||| Slovenia   (Kostanjšek & Kuntner, 2015) ||| Spain   (Branco et al., 2019) ||| Sweden   (Kronestedt, 2001) ||| Switzerland   (Maurer & Hänggi, 1990) ||| Turkey (Asia)   (Danışman et al., 2025a) ||| Ukraine   (Polchaninova & Prokopenko, 2019) ||| United Kingdom   (Lavery, 2019) |||
References

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Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510 pdf

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Bosmans R, Van Keer J, Russell-Smith A, Hadjiconstantis M, Komnenov M, Bosselaers J, Huber S, McCowan D, Snazell R, Decae A, Zoumides C, Kielhorn K-H, Oger P (2019c) Spiders of Cyprus (Araneae). A catalogue of all currently known species from Cyprus. Newsletter of the Belgian Arachnological Society 34, Suppl. 1: 1-173 pdf

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Christian S (2015) Micaria subopaca Westring, 1861, Scotophaeus blackwalli (Thorell, 1871) and S. scutulatus (L.Koch, 1866): three species of gnaphosids new to Luxembourg (Arachnida, Gnaophosidae). Bulletin de la Société des naturalistes luxembourgeois 117: 87-90 pdf

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Pfliegler W A, Schönhofer A, Niedbała W, Vella P, Sciberras A, Vella A (2017) New records of mites (Acari) and harvestmen (Opiliones) from Malta with a preliminary checklist of Maltese Arachnida. Soil Organisms 89: 85-110 pdf

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