Thanatus atratus Simon, 1875
Description
Male palp with small, rounded tegular apophysis. Embolus uniformly rounded. Median epigynal plate distinctly separating lateral pockets. Legs grey or light brown and spotted. Opisthosoma grey or light brown.
Body length male: 3.1-4.7 mmBody length female: 4.6-6.1 mm
Additional information
In dry and semi-dry steppe areas, on sea coasts and salt places, hibernation as juveniles
Distribution
Phenology
| Jan | Feb | Mar | Apr | May | Jun | Jul | Aug | Sep | Oct | Nov | Dec |
 |  |
Figures
Pedipalp
(Szita & Samu 2000)
Pedipalp, ventral view
(Kronestedt 1983)
Pedipalp, ventral view
(Pozzi & Hänggi 1998)
Pedipalp, ventral view
(Szita & Samu 2000)
Pedipalp
(Pozzi & Hänggi 1998)
Pedipalp
(Szita & Samu 2000)
Pedipalp, prolateral view
(Logunov 1996b)
Pedipalp, ventral view
(Logunov 1996b)
Pedipalp, retrolateral view
(Logunov 1996b)
Pedipalp, ventral view
(Muster & Thaler 2003)
Pedipalp, retrolateral view
(Muster & Thaler 2003)
Pedipalp, ventral view
(Hansen 1995)
Bulbus, apical view
(Logunov 1996b)
Tip of embolus, prolateral-distal view
(Muster & Thaler 2003)
Habitus
(Szita & Samu 2000)
Habitus, dorsal view
(Logunov 1996b)
Epigyne
(Pozzi & Hänggi 1998)
Epigyne
(Szita & Samu 2000)
Epigyne, ventral view
(Muster & Thaler 2003)
Epigyne, ventral view
(Logunov 1996b)
Epigyne
(Hansen 1995)
Preepigyne
(Řezáč & Růžička 2022)
Vulva
(Pozzi & Hänggi 1998)
Vulva
(Szita & Samu 2000)
Vulva, ventral view
(Muster & Thaler 2003)
Vulva
(Logunov 1996b)
Habitus
(Szita & Samu 2000)
Habitus, dorsal view
(Logunov 1996b)
Distribution List
"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.
References
Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129 
Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018)
Bosmans R, Van Keer J, Russell-Smith A, Kronestedt T, Alderweireldt M, Bosselaers J, De Koninck H (2013) Spiders of Crete (Araneae). A catalogue of all currently known species from the Greek island of Crete. Nieuwsbrief van de Belgische Arachnologische Vereniging 28 (Suppl. 1): 1-147
Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254
Danışman T, Kunt K B, Özkütük R S, Coşar İ (2024) The checklist of the spiders of Turkey. Version 2024 [last updated 01 January 2024], online at http://www.spidersofturkey.info
Grbac I, Katušić L, Lukić M (2019) Catalogue of spiders (Araneae) deposited in the Croatian Natural History Museum. Natura Croatica 28: 185-269
Grbić G, Hänggi A (2023) Additional spider records from Mt. Fruška Gora (Serbia) with a first insight into winter species. Acta Entomologica Serbica 27(2, 2022): 67-81
Hansen H (1995) Über die Arachniden-Fauna von urbanen Lebensräumen in Venedig - III. Die epigäischen Spinnen eines Stadtparkes (Arachnida: Araneae). Bollettino del Museo Civico di Storia Naturale di Venezia 44: 7-36
Komnenov M (2014) Spider fauna of the Osogovo Mt. Range, Northeastern Macedonia. Fauna Balkana 2: 1-267
Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91
Kronestedt T (1983) Spiders (Araneae) on the Great Alvar of the island of Öland, S Sweden. Entomologisk Tidskrift 104: 183-212
Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001]
Logunov D V (1996b) A critical review of the spider genera Apollophanes O. P.-Cambridge, 1898 and Thanatus C. L. Koch, 1837 in North Asia (Araneae, Philodromidae). Revue Arachnologique 11: 133-202
Muster C, Thaler K (2003) The Thanatus striatus species group in the eastern Alps, with description of Thanatus firmetorum sp. n. (Araneae: Philodromidae). Bulletin of the British Arachnological Society 12: 376-382
Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.
Pantini P, Isaia M (2019) Araneae.it: the online catalog of Italian spiders, with addenda on other arachnid orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). Fragmenta Entomologica 51: 127-152
Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App
Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp.
Pozzi S, Hänggi A (1998) Araignées nouvelles ou peu connues de la Suisse (Arachnida: Araneae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 71: 33-47
Řezáč M, Růžička V (2022) Thanatus aridorum Šilhavý, 1940 from Czechia is a junior synonym of Thanatus formicinus (Clerck, 1757) (Araneae: Philodromidae). Arachnologische Mitteilungen 63: 15-18
Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html
Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html
Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184
Szita É, Samu F (2000) Taxonomical review of Thanatus species (Philodromidae, Araneae) of Hungary. Acta Zoologica Academiae Scientiarum Hungaricae 46: 155-179
Van Keer J, Van Keer K, De Koninck H, Ramel G (2010) The arachnofauna (Araneae) of wetland Kerkini (Macedonia-Northern Central Greece). Acta Zoologica Bulgarica 62: 141-160
Vasiliev A I, Mikhailov K G (2023) Annotated checklist of spiders (Araneae) of the Republic of Moldova. Invertebrate Zoology 20: 307-342
Vrenozi B, Jäger P (2012) A faunistic study on ground-dwelling spiders (Araneae) in the Tirana district, Albania. Arachnologische Mitteilungen 44: 81-87
WSC (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, online at http://wsc.nmbe.ch (22.01.2024) doi: 10.24436/2
Zarikian N (2021) A survey of running crab spiders Philodromidae (Araneae) of Armenia. Bulletin of the Iraq Natural History Museum 16: 495-508