Theridiidae  Sundevall, 1833 / Heterotheridion  Wunderlich, 2008   1 species

Heterotheridion nigrovariegatum (Simon, 1873)

Beschreibung
Männchen

Pedipalpus Männchen: mit langer, geschwungener Tibia. Zeichnung bezüglich Randbreite an Prosoma und Sternum, Opisthosomaflecken und Beinringelung sehr variabel, mitunter das ganze Tier zitronengelb oder weiß. Prosoma gelblich-weiß, und Sternum gelblich-weiß mit feinem, dunklem Rand. Cheliceren gelblich. Beine weißlich gelb, schwarz geringelt. Opisthosoma ganz weiß oder gelblich mit großen schwarzen Flecken.

Körperlänge Männchen: 3.0-3.5 mm
Weibchen

Epigyne sehr klein, schwarz, zweiteilig. 

Körperlänge Weibchen: 3.0-4.1 mm
Zusätzliche Informationen

An sehr warmen Stellen auf Stauden und Gebüsch

Verbreitung
  Vorkommen
  kein Nachweis
Weltweite Verbreitung (WSC 2026): Europe, Turkey, Caucasus, Russia (Europe) to Central Asia, Iran, China
Phänologie
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Abbildungen
Verbreitungsnachweise
Albania   (Deltshev et al., 2011b) ||| Algeria   (Benhacene et al., 2023) ||| Austria   (Knoflach & Thaler, 1998) ||| Azerbaijan   (Otto, 2022) ||| Bulgaria   (Blagoev et al., 2018) ||| Croatia   (Milosevic, 2002) ||| Czechia   (Růžička & Řezáč, 2022a) ||| Egypt   (El-Hennawy, 2017a) ||| France   (Le Peru, 2007) ||| France / Corsica   (Le Peru, 2007) ||| Georgia   (Otto, 2022) ||| Germany   (Blick et al., 2016) ||| Greece   (Russell-Smith et al., 2011) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Isaia et al., 2025) ||| Italy / Sicily   (Isaia et al., 2025) ||| Libya   (Elkrew et al., 2024) ||| North Macedonia   (Komnenov, 2014) ||| Poland   (Rozwałka & Stanska, 2008) ||| Romania   (Weiss & Petrișor, 1999) ||| Russia, Central   (Ponomarev, 2022) ||| Russia, Eastern   (Esyunin & Efimik, 1996a) ||| Russia, Southern   (Otto, 2022; Ponomarev, 2022) ||| Serbia   (Grbić & Savić, 2010) ||| Slovakia   (Gajdoš et al., 1999) ||| Slovenia   (Kostanjšek & Kuntner, 2015) ||| Spain   (Branco et al., 2019) ||| Spain / Balearic Islands   (Branco et al., 2019; Barrientos & Febrer, 2018) ||| Switzerland   (Maurer & Hänggi, 1990) ||| Turkey (Asia)   (Danışman et al., 2025a) ||| Turkey (Europe)   (van Helsdingen, 2013; Demircan Aksan & Topçu, 2022c) ||| Ukraine   (Polchaninova & Prokopenko, 2019) |||
Literatur

Barrientos J A, Febrer J B (2018) Arañas (Arachnida, Araneae) de Menorca (Islas Baleares, España). 2: “adenda et corrigenda”. Descripción de tres especies nuevas. Revista Ibérica de Aracnología 33: 39-51 pdf

Benhacene R, Adjami A, Hadjeb A, Kermiche K, Ouakid M L (2023) Bibliographic checklist of the Algerian spider fauna (Araneae). Zootaxa 5352: 301-357 pdf

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510 pdf

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254 pdf

Danışman T, Kunt K B, Özkütük R S, Coşar İ (2025a) The checklist of the spiders of Turkey. Version 2025 [last updated 01 December 2025], online at http://www.spidersofturkey.info pdf

Deltshev C, Vrenozi B, Blagoev G, Lazarov S (2011b) Spiders of Albania – faunistic and zoogeographical review (Arachnida: Araneae). Acta Zoologica Bulgarica 63: 125-144 pdf

Demircan Aksan N, Topçu A (2022c) A preliminary checklist of the spider fauna of European Türkiye. Serket 19: 39-56 pdf

El-Hennawy H K (2017a) A list of Egyptian spiders (revised in 2017). Serket 15: 167-183 pdf

Elkrew H, Hamza A, Ghana S, Shaibi T, Swehli A I, Elmareme H, Ahmad A (2024) Checklist of the spiders (Araneae) of Libya. Pakistan Journal of Life and Social Sciences 22: 19346-19369 pdf

Esyunin S L, Efimik V E (1996a) Catalogue of the spiders (Arachnida, Aranei) of the Urals. KMK Scientific Press, Moscow, 228 pp. pdf

Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp. pdf

Grbić G, Savić D (2010) Contribution to the knowledge of the spider fauna (Arachnida, Araneae) on the Fruška Gora Mt. Acta Entomologica Serbica 15: 243-260 pdf

Helsdingen P J van (2013) A quick scan of the spider fauna of the European part of Turkey. Nieuwsbrief SPINED 33: 29-38 pdf

Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204 pdf

Jäger P, Staudt A, Schwarz B, Busse C (2000) Spinnen (Arachnida: Araneae) von Weinbergen und Weinbergsbrachen am Mittelrhein (Rheinland-Pfalz: Boppard, Oberwesel). Arachnologische Mitteilungen 19: 28-40 pdf

Knoflach B, Thaler K (1998) Kugelspinnen und verwandte Familien von Österreich: ökofaunistische Übersicht (Araneae: Theridiidae, Anapidae, Mysmenidae, Nesticidae). Stapfia 667-712 pdf

Komnenov M (2014) Spider fauna of the Osogovo Mt. Range, Northeastern Macedonia. Fauna Balkana 2: 1-267 pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Le Peru B (2007) Catalogue et répartition des araignées de France. Revue Arachnologique 16: 1-468 pdf

Maurer R, Hänggi A (1990) Katalog der schweizerischen Spinnen. Documenta Faunistica Helvetiae 12 pdf

Milosevic B (2002) Aranea – Popis vrsta – Checklist. Hrvatsko entomološko društvo – Inventar entomofaune Hrvatske [Aranea – List of species – Checklist. Croatian Entomological Society – Inventory of Croatian entomofauna]. pdf

Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.

Pesarini C (unpubl.)

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html pdf

Russell-Smith A, Allison R, Askins M, Blumsom W, Snazell R, Spilling C (2011) A provisional checklist and gazetteer of the spiders of Chios, Greece (Arachnida: Araneae). Bulletin of the British Arachnological Society 15: 133-167 pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Weiss I, Petrișor A (1999) List of the spiders (Arachnida: Araneae) from Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 41: 79-107 pdf

Wiehle H (1937) Spinnentiere oder Arachnoidea, VIII. 26: Familie Theridiidae oder Haubennetzspinnen (Kugelspinnen). Die Tierwelt Deutschlands 33: 119-222 pdf

WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (20.5.2026) doi: 10.24436/2 pdf

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