Gnaphosidae  Banks, 1892 / Gnaphosa  Latreille, 1804   71 species

Gnaphosa modestior Kulczyński, 1897

Description

Tibial apophysis twice as long as tibia. Prosoma black-brown. Femur and coxa shiny golden-yellow. Opisthosoma black-brown.

Body length male: 6.0-9.0 mm
Distribution
  Presence
  No data
Distribution comment: Turkey, Ukraine, Russia and Caucasus: see G. dolanskyi
Global distribution (WSC 2026): Italy, Austria, Czechia to Romania, Greece
Figures
Distribution List
Austria   (Řezáč et al., 2018a) ||| Azerbaijan  Removed (Nuruyeva & Zamani, 2025) ||| Croatia   (Milosevic, 2002) ||| Czechia   (Řezáč et al., 2021; Růžička & Řezáč, 2022a) ||| France  Removed (Danflous et al., 2020b) ||| Greece   (Pitta & Chatzaki, 2022) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Isaia et al., 2025) ||| Romania   (Weiss & Petrișor, 1999) ||| Serbia   (Stanković, 2023) ||| Slovakia   (Gajdoš et al., 1999) ||| Slovenia   (Kostanjšek & Kuntner, 2015) |||
References

Chyzer C, Kulczyński W (1897) Araneae Hungariae. Tomus II. Academia Scientarum Hungaricae, Budapest, pp. 147-366, pl. 6-10. pdf

Danflous S, Déjean S, Guerbaa K, Jacquet C, Lecigne S, Montagne D, Montardi Y, Villepoux O (2020b) De araneis Galliae IV. Actualisation du référentiel des araignées de France – première note. Espèces absentes de France ou mises en nomen dubium. Bulletin de l'Association Française d'Arachnologie 5: 10-32 pdf

Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp.

Grimm U (1985) Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF) 26: 1-318 pdf

Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204 pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Milosevic B (2002) Aranea – Popis vrsta – Checklist. Hrvatsko entomološko društvo – Inventar entomofaune Hrvatske [Aranea – List of species – Checklist. Croatian Entomological Society – Inventory of Croatian entomofauna]. pdf

Nuruyeva T, Zamani A (2025) An annotated checklist of the spiders (Arachnida: Araneae) of Azerbaijan. Zootaxa 5680: 1-168 pdf

Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.

Ovtsharenko V I, Platnick N I, Song D X (1992) A review of the North Asian ground spiders of the genus Gnaphosa (Araneae, Gnaphosidae). Bulletin of the American Museum of Natural History 212: 1-88 pdf

Pitta E, Chatzaki M (2022) Faunistic and zoogeographic analysis of sac and ground spiders (Cheiracanthiidae, Clubionidae and Gnaphosidae) in mainland Greece. Arachnologische Mitteilungen 64: 34-45 pdf

Řezáč M, Růžička V, Hula V, Dolanský J, Machač O, Roušar A (2021) Spiders newly observed in Czechia in recent years – overlooked or invasive species? BioInvasions Records 10: 555-566 pdf

Řezáč M, Růžička V, Oger P, Řezáčová V (2018a) European species of the Gnaphosa alpica complex (Araneae, Gnaphosidae). Zootaxa 4370: 289-294 pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Stanković B (2023) Diversity of spider fauna in the area of Jagodina (Serbia). In: Seydoşoğlu, S. & Durna Aydin, Ö (eds.) International Cukurova Agriculture and Veterinary Congress 4. Adana, pp. 590-615 pdf

Szmatona-Túri T, Kovács G, Vona-Túri D, Magos G (2019) Contribution to the knowledge on distribution in Hungary and habitat preferences of Gnaphosa modestior Kulczyński, 1897 a little-known spider species. Periodicum Biologorum 120: 135-139 pdf

Weiss I, Petrișor A (1999) List of the spiders (Arachnida: Araneae) from Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 41: 79-107 pdf

WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (20.5.2026) doi: 10.24436/2 pdf

Updates
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