Gnaphosa lucifuga (Walckenaer, 1802)
Description
Male palp lighter than body colour. Embolus robust, runs along ± the whole inner side of cymbium. Epigyne with very broad scapus, touching bar-like protrusions of lateral epigynal margins. Prosoma dark red-brown to black-brown, eye region darker. Chelicerae darker than prosoma. Legs lighter than prosoma. Opisthosoma dark red-brown to black-brown.
Body length male: 10.5-14.9 mmBody length female: 12.1-18.7 mm
Additional information
in sunny dry meadows, rock steppes and heathland, under stones during the day
Distribution
Phenology
| Jan | Feb | Mar | Apr | May | Jun | Jul | Aug | Sep | Oct | Nov | Dec |
 |  |
Figures
Pedipalp, ventral view
(Ovtsharenko et al. 1992)
Pedipalp, ventral view
(Ovtsharenko et al. 1992)
Pedipalp
(Ovtsharenko et al. 1992)
Pedipalp
(Roberts 1998)
Pedipalp, ventral view
(Tuneva & Esyunin 2003a)
Pedipalp, ventral view
(Løvbrekke unpubl.)
Pedipalp, ventral view
(Esyunin & Tuneva 2020)
Pedipalp, retrolateral view
(Taberlet unpubl.)
Pedipalp, ventral view
(Taberlet unpubl.)
Pedipalp, prolateral view
(Taberlet unpubl.)
Pedipalp, ventro-prolateral view
(Taberlet unpubl.)
Embolar base
(Esyunin & Tuneva 2020)
Palpal tibia, retrolateral view
(Løvbrekke unpubl.)
Palpal tibia, retrolateral view
(Esyunin & Tuneva 2020)
Epigyne
(Esyunin & Tuneva 2020)
Epigyne
(Ovtsharenko et al. 1992)
Epigyne
(Roberts 1998)
Epigyne
(Løvbrekke unpubl.)
Epigyne
(Esyunin & Tuneva 2020)
Epigyne
(Taberlet unpubl.)
Epigyne
(Taberlet unpubl.)
Scape, ventral view
(Løvbrekke unpubl.)
Vulva
(Ovtsharenko et al. 1992)
Vulva, dorsal view
(Tullgren 1946)
Vulva, ventral view
(Tullgren 1946)
Vulva
(Løvbrekke unpubl.)
Vulva, ventral view
(Taberlet unpubl.)
Vulva, dorsal view
(Taberlet unpubl.)
Vulva, posterior view
(Taberlet unpubl.)
Distribution List
"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.
References
Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129 
Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018)
Bosmans R, Van Keer K (2017) Een herziene soortenlijst van de Belgische spinnen (Araneae). Nieuwsbrief van de Belgische Arachnologische Vereniging 32: 39-69
Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254
Cera I (2018) The checklist of Latvian spiders (Arachnida: Araneae). Environmental and Experimental Biology 16: 139-152
Danışman T, Kunt K B, Özkütük R S, Coşar İ (2024) The checklist of the spiders of Turkey. Version 2024 [last updated 01 January 2024], online at http://www.spidersofturkey.info
Deltshev C, Komnenov M, Blagoev G, Georgiev T, Lazarov S, Stojkoska E, Naumova M (2013) Faunistic diversity of spiders (Araneae) in Galichitsa Mountain (FYR Macedonia). Biodiversity Data Journal 1(e977): 1-70
Deltshev C, Vrenozi B, Blagoev G, Lazarov S (2011b) Spiders of Albania – faunistic and zoogeographical review (Arachnida: Araneae). Acta Zoologica Bulgarica 63: 125-144
Esyunin S L, Tuneva T K (2020) A review of the family Gnaphosidae in the fauna of the Urals (Aranei), 6. Taxonomic remarks and new records, with description of a new species. Arthropoda Selecta 29: 103-120
Ivanov V V (2013) The checklist of Belarusian spiders (Arachnida, Araneae). Zoology and Ecology 4: 293-311
Komnenov M (2010) Checklist of spiders (Araneae) of Bosnia and Herzegovina. Uzizaž i Biospeld 5(2009): 52-69
Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91
Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001]
Løvbrekke H (unpubl.)
Milosevic B (2002) Aranea – Popis vrsta – Checklist. Hrvatsko entomološko društvo – Inventar entomofaune Hrvatske [Aranea – List of species – Checklist. Croatian Entomological Society – Inventory of Croatian entomofauna].
Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.
Ovtsharenko V I, Platnick N I, Song D X (1992) A review of the North Asian ground spiders of the genus Gnaphosa (Araneae, Gnaphosidae). Bulletin of the American Museum of Natural History 212: 1-88
Pantini P, Isaia M (2019) Araneae.it: the online catalog of Italian spiders, with addenda on other arachnid orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). Fragmenta Entomologica 51: 127-152
Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App
Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp.
Roberts M J (1998) Spinnengids. Tirion, Baarn, Netherlands, 397 pp.
Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html
Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html
Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184
Taberlet J-P (unpubl.)
Tullgren A (1946) Svensk spindelfauna. 3. Egentiliga spinddlar. Araneae Fam 5-7. Clubionidae, Zoridae och Gnaphosidae. Stockholm (Entomol Fören.): 141 pp.
Tuneva T K, Esyunin S L (2003a) A review of the Gnaphosidae fauna of the Urals (Aranei), 3. New species and new records, chiefly from the south Urals. Arthropoda Selecta 11: 223-234
Vasiliev A I, Mikhailov K G (2023) Annotated checklist of spiders (Araneae) of the Republic of Moldova. Invertebrate Zoology 20: 307-342
WSC (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, online at http://wsc.nmbe.ch (22.01.2024) doi: 10.24436/2