Male palp lighter than body colour. Embolus robust, runs along ± the whole inner side of cymbium. Epigyne with very broad scapus, touching bar-like protrusions of lateral epigynal margins. Prosoma dark red-brown to black-brown, eye region darker. Chelicerae darker than prosoma. Legs lighter than prosoma. Opisthosoma dark red-brown to black-brown.
Body length male: 10.5-14.9 mmin sunny dry meadows, rock steppes and heathland, under stones during the day
Jan | Feb | Mar | Apr | May | Jun | Jul | Aug | Sep | Oct | Nov | Dec |
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Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129
Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018)
Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). [N.B.: few records from this paper are not taken over and not commented] Zootaxa 4614: 201-254
Cera I (2018) The checklist of Latvian spiders (Arachnida: Araneae). Environmental and Experimental Biology 16: 139-152
Danışman T, Kunt K B, Özkütük R S (2019) The checklist of the spiders of Turkey. Version 2019 [last updated 1 June 2019], online at http://www.spidersofturkey.info
Deltshev C, Vrenozi B, Blagoev G, Lazarov S (2011b) Spiders of Albania – faunistic and zoogeographical review (Arachnida: Araneae). Acta Zoologica Bulgarica 63: 125-144
Esyunin S L, Tuneva T K (2020) A review of the family Gnaphosidae in the fauna of the Urals (Aranei), 6. Taxonomic remarks and new records, with description of a new species. Arthropoda Selecta 29: 103-120
Ivanov V V (2013) The checklist of Belarusian spiders (Arachnida, Araneae). Zoology and Ecology 4: 293-311
Komnenov M (2010) Checklist of spiders (Araneae) of Bosnia and Herzegovina. Uzizaž i Biospeld 5(2009): 52-69
Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91
Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001]
Løvbrekke H (unpubl.)
Otto S (2020) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 03.2020. Internet: caucasus-spiders.info.
Ovtsharenko V I, Platnick N I, Song D X (1992) A review of the North Asian ground spiders of the genus Gnaphosa (Araneae, Gnaphosidae). Bulletin of the American Museum of Natural History 212: 1-88
Pantini P, Isaia M (2019) Araneae.it: the online catalog of Italian spiders, with addenda on other arachnid orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). http://araneae.it. Fragmenta Entomologica 51: 127-152
Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App
Roberts M J (1998) Spinnengids. Tirion, Baarn, Netherlands, 397 pp.
Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184
Tullgren A (1946) Svensk spindelfauna. 3. Egentiliga spinddlar. Araneae Fam 5-7. Clubionidae, Zoridae och Gnaphosidae. Stockholm (Entomol Fören.): 141 pp.
Tuneva T K, Esyunin S L (2003a) A review of the Gnaphosidae fauna of the Urals (Aranei), 3. New species and new records, chiefly from the south Urals. Arthropoda Selecta 11: 223-234
WSC (2021) World Spider Catalog. Version 21.5. Natural History Museum Bern, online at http://wsc.nmbe.ch (03.01.2021) doi: 10.24436/2