Improphantes improbulus (Simon, 1929)
Description
Male palp with tapering lamella in distal third, pointed distally. Epigyne with strongly developed epigynal plates, enclosing scapus collar-like, therefore also visible from ventral, posterior part of scapus almost 4 times as wide as long. Prosoma yellow-brown, with or without dorsal spot. Legs clay-yellow. Opisthosoma grey-white to black.
Body length male: 2-2.3 mmBody length female: 2.1-2.7 mm
Additional information
in litter layer and small caves
Troglophile species (Mammola et al., 2022)
Character states from linyphiid key
Distribution
Phenology
| Jan | Feb | Mar | Apr | May | Jun | Jul | Aug | Sep | Oct | Nov | Dec |
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Figures
Pedipalp, ventral view
(Miller & Svatoň 1978)
Pedipalp, ventral view
(Heimer & Nentwig 1991)
Pedipalp, retrolateral view
(Thaler 1986c)
Pedipalp, prolateral view
(Thaler 1986c)
Paracymbium
(Thaler 1986c)
Epigyne
(Heimer 1978)
Epigyne
(Heimer & Nentwig 1991)
Epigyne
(Miller & Svatoň 1978)
Epigyne, lateral view
(Miller & Svatoň 1978)
Epigyne, antero-ventral view
(Thaler 1986c)
Epigyne/vulva, aboral view
(Thaler 1986c)
Epigyne, ventral view
(Thaler 1986c)
Vulva
(Miller & Svatoň 1978)
Distribution List
"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.
References
Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) 
Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510
Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254
Deltshev C, Vrenozi B, Blagoev G, Lazarov S (2011b) Spiders of Albania – faunistic and zoogeographical review (Arachnida: Araneae). Acta Zoologica Bulgarica 63: 125-144
Esyunin S L, Efimik V E (1996a) Catalogue of the spiders (Arachnida, Aranei) of the Urals. KMK Scientific Press, Moscow, 228 pp.
Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp.
Heimer S (1978) Zur intragenerischen Isolation der Arten der Gattung Pocadicnemis Simon 1884. Zoologische Abhandlungen, Staatliches Museum für Tierkunde Dresden 35: 101-112
Heimer S, Nentwig W (1991) Spinnen Mitteleuropas. Paul Parey Berlin
Komnenov M (2017) New data on spider fauna (Araneae) of Shar Mountain, north-western Macedonia. Proceedings of the 5th Congress of the Ecologists of Macedonia, with international participation (Ohrid, 19th-22nd October 2016). Special issues of the Macedonian Ecological Society 13: 44-61
Kreuels M, Staudt A, Christian S (2019) Die Spinnenfauna von Luxemburg – eine Zusammenstellung der Nachweise aus den Jahren 1906-2018 (Arachnida: Araneae). Bulletin de la Société des naturalistes luxembourgeois 121: 203-230
Le Peru B (2007) Catalogue et répartition des araignées de France. Revue Arachnologique 16: 1-468
Mammola S, Pavlek M, Huber B A, Isaia M, Ballarin F, Tolve M, Čupić I, Hesselberg T, Lunghi E, Mouron S, Graco-Roza C, Cardoso P (2022) A trait database and updated checklist for European subterranean spiders. Scientific Data 9(236): 1-13
Miller F, Svatoň J (1978) Einige seltene und bisher unbekannte Spinnenarten aus der Slowakei. Annotationes Zoologicae et Botanicae Bratislava 126: 1-19
Nae A (2010) Improphantes improbulus (Simon, 1929) (Araneae, Linyphiidae) new record for the Roumanian fauna. Travaux de l’Institut de Spéologie “Émile Racovitza” 49: 81-85
Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.
Pantini P, Isaia M (2019) Araneae.it: the online catalog of Italian spiders, with addenda on other arachnid orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). Fragmenta Entomologica 51: 127-152
Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp.
Růžička V (1996) Pavouci kamenitých sutí a pseudokrasových jeskyní Národního parku Podyjí [Spiders of stony debris and pseudokarst caves in Podyji National Park]. Příroda Sborník prací z ochrany přírody, Praha 3: 123-132
Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html
Staudt A, Weber D, Nährig D (2013) Webspinnen (Arachnida, Araneae) aus Höhlen des Großherzogtums Luxemburg. Ferrantia 69: 115-157
Thaler K (1986c) Vier bemerkenswerte Leptyphantes-Arten aus dem Mittelmeergebiet und aus Vorderasien (Arachnida: Aranei, Linyphiidae). Sitzungsberichte der Österreichischen Akademie der Wissenschaften (I) 194: 311-325
WSC (2025) World Spider Catalog. Version 26. Natural History Museum Bern, online at http://wsc.nmbe.ch (28.2.2025) doi: 10.24436/2