Dictynidae  O. Pickard-Cambridge, 1871 / Tolkienus  Cala-Riquelme, Crews & Esposito, 2025   2 species

Tolkienus ottoi (Marusik & Koponen, 2017)

Description
Distribution
  Presence
  No data
Global distribution (WSC 2026): Caucasus (Russia, Georgia, Azerbaijan), Iran, Kazakhstan
Figures
Distribution List
Azerbaijan   (Otto, 2022) ||| Georgia   (Otto, 2022) ||| Russia, Southern   (Otto, 2022; Ponomarev, 2022) |||
References

Marusik Y M, Koponen S (2017) On two sibling species of Dictyna (Araneae: Dictynidae) from Ukraine and Caucasus. Entomologica Fennica 28: 41-48 pdf

Nekhaeva A, Kim L, Ishayeva A, Ozernoy A, Sozontov A (2026) Spider (Arachnida, Araneae) fauna of the lowland part of the Balkhash-Alakol Basin (SE Kazakhstan): an integrated assessment. Biodiversity Data Journal 14(e181501): 1-40 pdf

Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.

Otto S, Japoshvili G (2018) The spiders (Arachnida: Araneae) of the Lagodekhi Reserve, Georgia: faunistic results of a transect study and an updated checklist. Arachnology 17: 375-391 pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Ponomarev A V, Aliev M A, Khabiev G N, Shmatko V Y (2019a) New data on the spider fauna (Aranei) of Dagestan, Russia. Arthropoda Selecta 28: 309-334 pdf

WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (20.5.2026) doi: 10.24436/2 pdf

Zamani A, Nadolny A A, Esyunin S L, Marusik Y M (2021) New data on the spider fauna of Iran (Arachnida: Araneae), part VIII. Zoosystematica Rossica 30: 279-297 pdf

Updates
21-02-2026 Image insert
04-12-2021 Image insert
04-07-2019 Image insert
01-07-2019 Distribution update Detail
24-07-2018 Image insert
23-07-2018 Distribution update Detail
07-03-2017 Image insert
07-03-2017 Distribution update Detail