Gnaphosa bicolor (Hahn, 1833)
Description
Palp female: yellow-orange, male: with short, stout spines on femur and tibia. Epigyne with striking transverse folds of epigynal groove. Prosoma dark red-brown to black-brown. Femur yellow-orange. Metatarsus and coxa yellow-orange. Opisthosoma dark red-brown to black-brown.
Body length male: 6.2-8.4 mmBody length female: 6.8-10.0 mm
Additional information
in hilly and mountainous terrain, in bright forests or rocky steppes, under stones during the day, in leaf litter layer, up to 1000 m
Distribution
Phenology
| Jan | Feb | Mar | Apr | May | Jun | Jul | Aug | Sep | Oct | Nov | Dec |
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Figures
Pedipalp, ventral view
(Ovtsharenko et al. 1992)
Pedipalp
(Ovtsharenko et al. 1992)
Pedipalp, ventral view
(Løvbrekke unpubl.)
Pedipalp, retrolateral view
(Grimm 1985)
Pedipalp, ventral view
(Grimm 1985)
Pedipalp, retrolateral view
(Almquist 2007)
Pedipalp, ventral view
(Almquist 2007)
Palpal tibia, retrolateral view
(Løvbrekke unpubl.)
Palpal patella and femur, retrolateral view
(Grimm 1985)
Palpal tibia, dorsal view
(Almquist 2007)
Epigyne
(Ovtsharenko et al. 1992)
Epigyne
(Løvbrekke unpubl.)
Epigyne
(Grimm 1985)
Epigyne
(Almquist 2007)
Vulva
(Ovtsharenko et al. 1992)
Vulva, ventral view
(Tullgren 1946)
Vulva
(Løvbrekke unpubl.)
Vulva, dorsal view
(Grimm 1985)
Vulva, dorsal view
(Almquist 2007)
Habitus
(Tullgren 1946)
Distribution List
"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.
References
Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129 
Almquist S (2007) Swedish Araneae, part 2 – families Dictynidae to Salticidae. Insect Systematics & Evolution, Supplement 63(2006): 285-601
Arnold K (2001) Beitrag zur Spinnenfauna (Arachnida, Araneae) des Fürstentums Liechtenstein. Berichte der Botanisch-Zoologischen Gesellschaft Liechtenstein-Sargans-Werdenberg 28: 211-244
Biteniekytė M, Rėlys V (2012) The checklist of Lithuanian spiders (Arachnida: Araneae). Biologija 57: 148-158
Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018)
Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510
Bosmans R, Chatzaki M (2005) A catalogue of the spiders of Greece – a critical review of all spider species cited from Greece with their localities. Nieuwsbrief van de Belgische arachnologische Vereniging 20 (2, suppl.): 1-124
Cera I (2018) The checklist of Latvian spiders (Arachnida: Araneae). Environmental and Experimental Biology 16: 139-152
Danışman T, Kunt K B, Özkütük R S, Coşar İ (2025a) The checklist of the spiders of Turkey. Version 2025 [last updated 01 December 2025], online at http://www.spidersofturkey.info
Deltshev C D, Ćurčić B P M, Blagoev GA (2003b) The spiders of Serbia. Inst. Zoology, Fac. Biology, Belgrade & Inst. Zoology, Bulg. Acad. Sci., Sofia. 832 pp.
Esyunin S L, Efimik V E (1996a) Catalogue of the spiders (Arachnida, Aranei) of the Urals. KMK Scientific Press, Moscow, 228 pp.
Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp.
Grimm U (1985) Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF) 26: 1-318
Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204
Ivanov V V (2013) The checklist of Belarusian spiders (Arachnida, Araneae). Zoology and Ecology 4: 293-311
Komnenov M (2014) Spider fauna of the Osogovo Mt. Range, Northeastern Macedonia. Fauna Balkana 2: 1-267
Koponen S, Fritzén N R, Pajunen T (2016) Checklist of spiders in Finland (Araneae). 6th version, December 2016, online at http://biolcoll.utu.fi/arach/checklist_of_spiders_in_Finland.htm
Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91
Kreuels M, Staudt A, Christian S (2019) Die Spinnenfauna von Luxemburg – eine Zusammenstellung der Nachweise aus den Jahren 1906-2018 (Arachnida: Araneae). Bulletin de la Société des naturalistes luxembourgeois 121: 203-230
Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001]
Le Peru B (2007) Catalogue et répartition des araignées de France. Revue Arachnologique 16: 1-468
Løvbrekke H (unpubl.)
Maurer R, Hänggi A (1990) Katalog der schweizerischen Spinnen. Documenta Faunistica Helvetiae 12
Milosevic B (2002) Aranea – Popis vrsta – Checklist. Hrvatsko entomološko društvo – Inventar entomofaune Hrvatske [Aranea – List of species – Checklist. Croatian Entomological Society – Inventory of Croatian entomofauna].
Naumova M, Hristovski S, Hristov G H (2016b) Spiders (Arachnida: Araneae) from Prespa National Park, Albania. Acta Zoologica Bulgarica 68: 503-511
Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.
Ovtsharenko V I, Platnick N I, Song D X (1992) A review of the North Asian ground spiders of the genus Gnaphosa (Araneae, Gnaphosidae). Bulletin of the American Museum of Natural History 212: 1-88
Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App
Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp.
Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html
Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html
Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184
Scharff N, Gudik-Sørensen O (2006) Katalog over Danmarks edderkopper (Araneae). Catalogue of the spiders of Denmark (Araneae). Entomologiske Meddelelser 74: 3-71
Thaler K, Knoflach B (2004b) Zur Faunistik der Spinnen (Araneae) von Österreich: Gnaphosidae, Thomisidae (Dionycha pro parte). Linzer Biologische Beiträge 36: 417-484
Tullgren A (1946) Svensk spindelfauna. 3. Egentiliga spinddlar. Araneae Fam 5-7. Clubionidae, Zoridae och Gnaphosidae. Stockholm (Entomol Fören.): 141 pp.
Vilbaste A (1987) Eesti Ämblikud (Aranei). Annoteeritud nimestik. Estonian spiders (Aranei). An annotated check list. Valgus, Tallinn. 113 pp. & 512 maps on 67 unpag. pages
Weiss I, Petrișor A (1999) List of the spiders (Arachnida: Araneae) from Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 41: 79-107
WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (13.2.2026) doi: 10.24436/2
Updates
| 03-02-2026 | Distribution insert | Detail |
| 15-12-2025 | Image insert | |
| 24-07-2025 | Distribution update | Detail |
| 14-01-2025 | Image insert | |
| 15-01-2023 | Distribution update | Detail |
| 28-06-2020 | Image insert | |
| 14-05-2020 | Distribution insert | Detail |
| 14-10-2019 | Image insert | |
| 11-05-2018 | Distribution update | Detail |
| 01-12-2013 | Distribution update | Detail |
| 09-04-2013 | Distribution update | Detail |