Linyphiidae  Blackwall, 1859 / Theonina  Simon, 1929   4 species

Theonina cornix (Simon, 1881)

Description
Male

Cymbium with 2 robust processes on base. Embolus thick and bent.
Prosoma dark brown to black. Sternum dark brown. Chelicerae brown, relatively long. Legs yellow-brown, thin and short. Opisthosoma yellow-brown to black.

Body length male: 1.2-1.5 mm
Female

Epigyne with scapus, longer than wide. 

Body length female: 1.5 mm
Additional information

In dry forests and on shrubs

Character states from linyphiid key
Distribution
  Presence
  No data
Distribution comment: Georgia: cf.
Global distribution (WSC 2026): Europe, North Africa, Georgia?
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List

"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.

Algeria   (Benhacene et al., 2023) ||| Austria   (Thaler, 1995) ||| Belgium   (Bosmans & Van Keer, 2017) ||| Bosnia and Herzegovina   (Komnenov, 2010) ||| Czechia   (Růžička & Řezáč, 2022a) ||| France   (Le Peru, 2007) ||| Georgia   (Seropian et al., 2023b) ||| Germany   (Blick et al., 2016) ||| Greece / Crete   (Bosmans et al., 2013) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Isaia et al., 2025) ||| Italy / Sardinia   (Isaia et al., 2025; Bosmans & Colombo, 2015) ||| Morocco   (Benhalima & Bosmans, 2024) ||| Netherlands   (van Helsdingen, 2025a; van Helsdingen et al., 2013) ||| North Macedonia   (Komnenov, 2014) ||| Portugal   (Branco et al., 2019) ||| Romania   (Weiss & Petrișor, 1999) ||| Russia, Central   (no references) ||| Slovakia   (Gajdoš et al., 1999) ||| Slovenia   (Kostanjšek & Kuntner, 2015) ||| Spain   (Branco et al., 2019) ||| Switzerland   (Maurer & Hänggi, 1990) ||| Tunisia   (Kmira et al., 2025) |||
References

Benhacene R, Adjami A, Hadjeb A, Kermiche K, Ouakid M L (2023) Bibliographic checklist of the Algerian spider fauna (Araneae). Zootaxa 5352: 301-357 pdf

Benhalima S, Bosmans R (2024) First historical checklist of spiders (Arachnida: Araneae) from Morocco, between 1840–2024. Zootaxa 5555: 407-535 pdf

Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510 pdf

Bosmans R (2006a) Contribution to the knowledge of the Linyphiidae of the Maghreb. Part XI. Miscellaneous linyphiid genera and additions (Araneae: Linyphiidae: Linyphiinae). Bulletin & Annales de la Société Entomologique de Belgique 141: 125-161 pdf

Bosmans R, Colombo M (2015) New species of spiders from Sardinia (Araneae), with ecological notes on Lipocrea epeiroides (O. Pickard-Cambridge, 1872) (Araneae: Araneidae). Arachnology 16: 319-332 pdf

Bosmans R, Van Keer J, Russell-Smith A, Kronestedt T, Alderweireldt M, Bosselaers J, De Koninck H (2013) Spiders of Crete (Araneae). A catalogue of all currently known species from the Greek island of Crete. Nieuwsbrief van de Belgische Arachnologische Vereniging 28 (Suppl. 1): 1-147 pdf

Bosmans R, Van Keer K (2017) Een herziene soortenlijst van de Belgische spinnen (Araneae). Nieuwsbrief van de Belgische Arachnologische Vereniging 32: 39-69 pdf

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254 pdf

Deeleman-Reinhold C L (1978b) Revision of the cave-dwelling and related spiders of the genus Troglohyphantes Joseph (Linyphiidae), with special reference to the yugoslav species. Slovenska Akademija Znanosti in Umetnosti, Razred za Prirodoslovne Vede, Classis IV, Historia Naturalis (Prirod. Vede) 23: 1-220 pdf

Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp.

Helsdingen P J van (2025a) Catalogus van de Nederlandse spinnen. Versie 2025.1. 249 pp. pdf

Helsdingen P J van, IJland S, Noordijk J, Wijnhoven H (2013) Arachniden van het Roerdal: 1000-soortendag met enkele aanvullingen (Arachnida: Araneae, Opiliones, Pseudoscorpiones). Nieuwsbrief SPINED 33: 21-28 pdf

Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204 pdf

Kmira G, Nasri-Ammar, Petillon J, Bosmans R (2025) Catalog of the spiders of Tunisia (Arachnida: Araneae). Zootaxa 5641: 1-95 pdf

Komnenov M (2010) Checklist of spiders (Araneae) of Bosnia and Herzegovina. Uzizaž i Biospeld 5(2009): 52-69 pdf

Komnenov M (2014) Spider fauna of the Osogovo Mt. Range, Northeastern Macedonia. Fauna Balkana 2: 1-267 pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Le Peru B (2007) Catalogue et répartition des araignées de France. Revue Arachnologique 16: 1-468 pdf

Maurer R, Hänggi A (1990) Katalog der schweizerischen Spinnen. Documenta Faunistica Helvetiae 12 pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Saaristo M I (1974) Taxonomical analysis of Theonina cornix (Simon 1881), the type-species of the genus Theonina Simon, 1929 (Araneae, Linyphiidae). Annales Zoologici Fennici 11: 240-243 pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Seropian A, Bulbulashvili N, Otto S, Krammer H-J, Kachlishvili N, Datunashvili A (2023b) Picking pearls from the Silk Road: insights into the spider (Arachnida, Araneae) diversity in Georgia from the Caucasus barcode of life project. Part II. Caucasiana 2: 231-297 pdf

Thaler K (1995) Beiträge zur Spinnenfauna von Nordtirol - 5. Linyphiidae 1: Linyphiinae (sensu Wiehle) (Arachnida: Araneida). Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck 82: 153-190 pdf

Weiss I, Petrișor A (1999) List of the spiders (Arachnida: Araneae) from Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 41: 79-107 pdf

Wiehle H (1956) Spinnentiere oder Arachnoidea, X. 28: Familie Linyphiidae-Baldachinspinnen. Die Tierwelt Deutschlands 44: 1-337 pdf

WSC (2025) World Spider Catalog. Version 26. Natural History Museum Bern, online at http://wsc.nmbe.ch (28.2.2025) doi: 10.24436/2 pdf

Updates
21-10-2025 Distribution update Detail
13-02-2025 Distribution update Detail
14-12-2023 Distribution insert Detail
06-08-2020 Image insert
08-06-2020 Distribution update Detail
13-01-2016 Distribution update Detail
24-09-2015 Distribution update Detail
15-10-2013 Distribution update Detail