Philodromidae  Thorell, 1869 / Thanatus  C. L. Koch, 1837   37 species

Thanatus formicinus  (Clerck, 1757)

Description

Tibial apophysis situated retrolaterally, with broad triangular base. Epigyne with round, sclerotised copulatory opening. Prosoma light brown to light reddish brown. Legs light brown, to light reddish brown. Opisthosoma light brown to light reddish brown.

Body length male: 5.2-7.4 mm
Body length female: 6.9-12 mm
Additional information

In dry and semi-dry meadows and xerothermic forest steppes

Distribution
  Presence
  No data
Distribution comment: Svalbard: introduced (Coulson et al. 2014).
Global distribution (WSC 2024): North America, Europe, North Africa, Turkey, Caucasus, Russia (Europe to Far East), Iraq, Iran, Kazakhstan, Central Asia, China, Japan
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Overwinter as subadults or adults, i.e. adults appear early in the year (March)
Figures
Distribution List

"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.

Albania   (Kůrka et al., 2020) ||| Algeria   (Benhacene et al., 2023) ||| Andorra   (Branco et al., 2019) ||| Armenia   (Zarikian, 2021) ||| Austria   (no references) ||| Azerbaijan  Removed (Logunov & Huseynov, 2008) ||| Belarus   (Ivanov, 2013) ||| Belgium   (Bosmans & Van Keer, 2017) ||| Bulgaria   (Blagoev et al., 2018) ||| Croatia   (Milosevic, 2002) ||| Czechia   (Růžička & Řezáč, 2022a) ||| Denmark   (no references) ||| Estonia   (no references) ||| Finland   (Koponen et al., 2016) ||| France   (no references) ||| France / Corsica   (Le Peru, 2007) ||| Georgia   (Otto, 2022) ||| Germany   (no references) ||| Greece   (no references) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Pantini & Isaia, 2019) ||| Italy / Sardinia   (Pantini & Isaia, 2019) ||| Kosovo   (Geci et al., 2023) ||| Latvia   (Cera, 2018) ||| Liechtenstein   (Arnold, 2001) ||| Lithuania   (no references) ||| Luxembourg   (Hermann, 1998; Kreuels et al., 2019) ||| Moldova   (Vasiliev & Mikhailov, 2023) ||| Netherlands   (no references) ||| North Macedonia   (Komnenov, 2014) ||| Norway   (Aakra & Hauge, 2003) ||| Poland   (Rozwałka & Stanska, 2008) ||| Portugal   (Branco et al., 2019) ||| Romania   (no references) ||| Russia, Central   (Ponomarev, 2022) ||| Russia, Eastern   (no references) ||| Russia, Northern   (Tanasevitch & Kamayev, 2011) ||| Russia, Southern   (Otto, 2022; Martynovchenko & Mikhailov, 2014; Ponomarev, 2022) ||| Russia, Western   (no references) ||| Serbia   (no references) ||| Slovakia   (no references) ||| Slovenia   (Kostanjšek & Kuntner, 2015) ||| Spain   (Branco et al., 2019) ||| Svalbard   (Coulson et al., 2014) ||| Sweden   (Kronestedt, 2001) ||| Switzerland   (no references) ||| Turkey (Asia)   (Danışman et al., 2024a) ||| Ukraine   (Polchaninova & Prokopenko, 2019) ||| United Kingdom   (Lavery, 2019) |||
References

Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129 pdf

Arnold K (2001) Beitrag zur Spinnenfauna (Arachnida, Araneae) des Fürstentums Liechtenstein. Berichte der Botanisch-Zoologischen Gesellschaft Liechtenstein-Sargans-Werdenberg 28: 211-244 pdf

Azheganova N S (1968) Kratkii opredelitel' paukov (Aranei) lesnoi i lesostepnoi zony SSSR. Akademia Nauk SSSR, 149pp. pdf

Barrientos J A, Urones M C (1985) La colección de araneidos del Departamento de Zoología de la Universidad de Salamanca, V: arañas clubionoideas y tomisoideas. Boletín de la Asociación Española de Entomología 9: 349-366 pdf

Benhacene R, Adjami A, Hadjeb A, Kermiche K, Ouakid M L (2023) Bibliographic checklist of the Algerian spider fauna (Araneae). Zootaxa 5352: 301-357 pdf

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Bosmans R, Van Keer K (2017) Een herziene soortenlijst van de Belgische spinnen (Araneae). Nieuwsbrief van de Belgische Arachnologische Vereniging 32: 39-69 pdf

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254 pdf

Cera I (2018) The checklist of Latvian spiders (Arachnida: Araneae). Environmental and Experimental Biology 16: 139-152 pdf

Coulson S J et al. (2014) The terrestrial and freshwater invertebrate biodiversity of the archipelagoes of the Barents Sea; Svalbard, Franz Josef Land and Novaya Zemlya. Soil Biology & Biochemistry 68: 440-470 pdf

Danışman T, Kunt K B, Özkütük R S, Coşar İ (2024a) The checklist of the spiders of Turkey. Version 2024 [last updated 01 January 2024], online at http://www.spidersofturkey.info pdf

Geci D, Naumova M, Ibrahimi H, Grapci-Kotori L, Gashi A, Bilalli A, Musliu M (2023) Contribution to spider fauna (Arachnida: Araneae) from Bjeshkët e Nemuna mountains (Kosovo). Natura Croatica 32: 525-547 pdf

Hermann E (1998) Die Spinnen (Araneae) ausgewählter Halbtrockenrasen im Osten Luxemburgs. Bulletin de la Société des naturalistes Luxembourgeois 99: 189-199 pdf

Ivanov V V (2013) The checklist of Belarusian spiders (Arachnida, Araneae). Zoology and Ecology 4: 293-311 pdf

Komnenov M (2014) Spider fauna of the Osogovo Mt. Range, Northeastern Macedonia. Fauna Balkana 2: 1-267 pdf

Koponen S, Fritzén N R, Pajunen T (2016) Checklist of spiders in Finland (Araneae). 6th version, December 2016, online at http://biolcoll.utu.fi/arach/checklist_of_spiders_in_Finland.htm pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Kreuels M, Staudt A, Christian S (2019) Die Spinnenfauna von Luxemburg – eine Zusammenstellung der Nachweise aus den Jahren 1906-2018 (Arachnida: Araneae). Bulletin de la Société des naturalistes luxembourgeois 121: 203-230 pdf

Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001] pdf

Kůrka A, Naumova M, Indzhov S, Deltshev C (2020) New faunistic and taxonomic data on the spider fauna of Albania (Arachnida: Araneae). Arachnologische Mitteilungen 59: 8-21 pdf

Lavery A (2019) A revised checklist of the spiders of Great Britain and Ireland. Arachnology 18: 196-212 pdf

Le Peru B (2007) Catalogue et répartition des araignées de France. Revue Arachnologique 16: 1-468 pdf

Locket G H, Millidge A F (1951) British spiders. Vol. I. Ray Society, London, 310 pp. pdf

Logunov D V (1996b) A critical review of the spider genera Apollophanes O. P.-Cambridge, 1898 and Thanatus C. L. Koch, 1837 in North Asia (Araneae, Philodromidae). Revue Arachnologique 11: 133-202 pdf

Logunov D V, Huseynov E F (2008) A faunistic review of the spider family Philodromidae (Aranei) of Azerbaijan. Arthropoda Selecta 17: 117-131 pdf

Løvbrekke H (unpubl.)

Martynovchenko F A, Mikhailov K G (2014) Spiders (Aranei) of Teberda State Reserve: fauna and biotopic distribution. Euroasian Entomological Journal 13: 355-371 pdf

Milosevic B (2002) Aranea – Popis vrsta – Checklist. Hrvatsko entomološko društvo – Inventar entomofaune Hrvatske [Aranea – List of species – Checklist. Croatian Entomological Society – Inventory of Croatian entomofauna]. pdf

Morka G H (unpubl.)

Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.

Pantini P, Isaia M (2019) Araneae.it: the online catalog of Italian spiders, with addenda on other arachnid orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). Fragmenta Entomologica 51: 127-152 pdf

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Řezáč M, Růžička V (2022) Thanatus aridorum Šilhavý, 1940 from Czechia is a junior synonym of Thanatus formicinus (Clerck, 1757) (Araneae: Philodromidae). Arachnologische Mitteilungen 63: 15-18 pdf

Roberts M J (1995) Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp. pdf

Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Šilhavý V (1940) Thanatus aridorum n. sp. novy pavouk z Mohelnske hadcove stepi. Folia Entomologica 3: 14-15 pdf

Szita É, Samu F (2000) Taxonomical review of Thanatus species (Philodromidae, Araneae) of Hungary. Acta Zoologica Academiae Scientiarum Hungaricae 46: 155-179 pdf

Tanasevitch A V, Kamayev I O (2011) Spiders of Kola Peninsula, Russia (Arachnida: Aranei). Caucasian Entomological Bulletin 7: 7-32 pdf

Urones C (1986) La familia Philodromidae (Araneae) en el centro-oeste de la Península Ibérica. Boletín de la Asociación Española de Entomología 10: 231-244 pdf

Vasiliev A I, Mikhailov K G (2023) Annotated checklist of spiders (Araneae) of the Republic of Moldova. Invertebrate Zoology 20: 307-342 pdf

WSC (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, online at http://wsc.nmbe.ch (22.01.2024) doi: 10.24436/2 pdf

Zarikian N (2021) A survey of running crab spiders Philodromidae (Araneae) of Armenia. Bulletin of the Iraq Natural History Museum 16: 495-508 pdf

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