Theridiidae  Sundevall, 1833 / Enoplognatha  Pavesi, 1880   30 species

Enoplognatha quadripunctata  Simon, 1884

Description
Male

Prosoma 0.85-1.9 mm long. Chelicerae as in E. thoracica: one large tooth with denticles at its base, and one smaller tooth.

Body length male: 1.8-3.8 mm
Female

Epigyne not strongly sclerotized, median part flat and trapezoid. Prosoma 1-1.6 mm long. Colouration as in E. thoracica, usually with opisthosomal spots, but sometimes uniformly coloured.

Body length female: 2.2-4.1 mm
Additional information

Usually in dry habitats, sand dunes, maquis, pine forests, stony hillsides, dry shrubs, also in chestnut woods, grasslands and marshes.

Distribution
  Presence
  No data
Global distribution (WSC 2020): Mediterranean, Caucasus, Kazakhstan
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List
Albania   (Deltshev et al., 2011b) ||| Algeria   (Bosmans, unpubl.) ||| Azerbaijan   (Otto, 2020) ||| Bulgaria   (Blagoev et al., 2018) ||| Cyprus   (Bosmans et al., 2019c) ||| France / Corsica   (no references) ||| Georgia   (Otto, 2020) ||| Greece   (no references) ||| Greece / Crete   (no references) ||| Morocco   (Bosmans, unpubl.) ||| North Macedonia   (Komnenov, 2014) ||| Portugal   (Branco et al., 2019) ||| Russia, Southern   (Otto, 2020) ||| Spain   (Branco et al., 2019) ||| Spain / Balearic Islands   (Branco et al., 2019) ||| Tunisia   (Bosmans, unpubl.) ||| Turkey (Asia)   (Danışman et al., 2019) ||| Turkey (Europe)   (Danışman et al., 2019) |||
References

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Bosmans R, Van Keer J (1999) The genus Enoplognatha Pavesi, 1880 in the Mediterranean region (Araneae: Theridiidae). Bulletin of the British Arachnological Society 11: 209-241 pdf

Bosmans R, Van Keer J, Russell-Smith A, Hadjiconstantis M, Komnenov M, Bosselaers J, Huber S, McCowan D, Snazell R, Decae A, Zoumides C, Kielhorn K-H, Oger P (2019c) Spiders of Cyprus (Araneae). A catalogue of all currently known species from Cyprus. Newsletter of the Belgian Arachnological Society 34, Suppl. 1: 1-173 pdf

Bosmans R. (unpubl.) Provisional list of spiders of North Africa. Database excerpt Aug. 2019

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae).
[N.B.: few records from this paper are not taken over and not commented] Zootaxa 4614: 201-254 pdf

Danışman T, Kunt K B, Özkütük R S (2019) The checklist of the spiders of Turkey. Version 2019 [last updated 1 June 2019], online at http://www.spidersofturkey.info pdf

Deltshev C, Vrenozi B, Blagoev G, Lazarov S (2011b) Spiders of Albania – faunistic and zoogeographical review (Arachnida: Araneae). Acta Zoologica Bulgarica 63: 125-144 pdf

Komnenov M (2014) Spider fauna of the Osogovo Mt. Range, Northeastern Macedonia. Fauna Balkana 2: 1-267 pdf

Le Peru B (2011) The spiders of Europe, a synthesis of data: Volume 1 Atypidae to Theridiidae. Mémoires de la Société Linnéenne de Lyon 2: 1-522 pdf

Otto S (2020) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 03.2020. Internet: caucasus-spiders.info.

World Spider Catalog (2020) World Spider Catalog. Version 21.0. Natural History Museum Bern, online at http://wsc.nmbe.ch (20.03.2020) doi: 10.24436/2 pdf

Updates
21-10-2015 Distribution update Detail
04-12-2013 Distribution update Detail