Zelotes clivicola (L. Koch, 1870)
Description
Male
Embolus long and pointed. Keel at inner side of bulbus covered by distal part of bulbus. Prosoma black-brown to black. Opisthosoma black-brown to black, with dark brown scuticula.
Body length male: 4.3-5.5 mmFemale
Epigyne with only slightly rounded cuticular fold.
Body length female: 4.5-7.1 mmAdditional information
In open forests, dwarf shrub heath and bogs, up to 2000 m.
Similar to Z. gallicus.
Distribution
Phenology
| Jan | Feb | Mar | Apr | May | Jun | Jul | Aug | Sep | Oct | Nov | Dec |
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Figures
Pedipalp, retrolateral view
(Thaler 1989)
Pedipalp, retrolateral view
(Tullgren 1946)
Pedipalp, ventral view
(Tullgren 1946)
Pedipalp, ventral view
(Løvbrekke unpubl.)
Pedipalp, ventral view
(Løvbrekke unpubl.)
Pedipalp, ventral view
(Kostanjšek unpubl.)
Pedipalp, ventral view
(Kostanjšek unpubl.)
Pedipalp, retrolateral view
(Grimm 1985)
Pedipalp, ventral view
(Grimm 1985)
Pedipalp, ventral view
(Almquist 2007)
Pedipalp, retrolateral view
(Almquist 2007)
Tip of pedipalp, ventral view
(Thaler 1989)
Bulbus, ventral view
(Miller 1967)
Embolus, dorsal view
(Løvbrekke unpubl.)
Embolus, ventral view
(Løvbrekke unpubl.)
Embolic division
(Miller 1967)
Palpal tibia, dorsal view
(Almquist 2007)
Epigyne
(Brændegaard 1966)
Epigyne, ventral view
(Tullgren 1946)
Epigyne, ventral view
(Izmailova 1989)
Epigyne, ventral view
(Løvbrekke unpubl.)
Epigyne, ventral view
(Løvbrekke unpubl.)
Epigyne
(Grimm 1985)
Epigyne
(Grimm 1985)
Epigyne
(Morka unpubl.)
Epigyne
(Almquist 2007)
Vulva
(Jézéquel 1962a)
Vulva, dorsal view
(Tullgren 1946)
Vulva, dorsal view
(Løvbrekke unpubl.)
Vulva, dorsal view
(Grimm 1985)
Vulva, dorsal view
(Almquist 2007)
Distribution List
"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.
References
Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129 
Almquist S (2007) Swedish Araneae, part 2 – families Dictynidae to Salticidae. Insect Systematics & Evolution, Supplement 63(2006): 285-601
Arnold K (2001) Beitrag zur Spinnenfauna (Arachnida, Araneae) des Fürstentums Liechtenstein. Berichte der Botanisch-Zoologischen Gesellschaft Liechtenstein-Sargans-Werdenberg 28: 211-244
Biteniekytė M, Rėlys V (2012) The checklist of Lithuanian spiders (Arachnida: Araneae). Biologija 57: 148-158
Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018)
Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510
Bosmans R, Chatzaki M (2005) A catalogue of the spiders of Greece – a critical review of all spider species cited from Greece with their localities. Nieuwsbrief van de Belgische arachnologische Vereniging 20 (2, suppl.): 1-124
Bosmans R, Van Keer J, Russell-Smith A, Kronestedt T, Alderweireldt M, Bosselaers J, De Koninck H (2013) Spiders of Crete (Araneae). A catalogue of all currently known species from the Greek island of Crete. Nieuwsbrief van de Belgische Arachnologische Vereniging 28 (Suppl. 1): 1-147
Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254
Brændegaard J (1966) Edderkopper: Eller Spindlere I Danmarks Fauna 72: 1-224
Cera I (2018) The checklist of Latvian spiders (Arachnida: Araneae). Environmental and Experimental Biology 16: 139-152
Danışman T, Kunt K B, Özkütük R S, Coşar İ (2025a) The checklist of the spiders of Turkey. Version 2025 [last updated 01 December 2025], online at http://www.spidersofturkey.info
Deltshev C D, Ćurčić B P M, Blagoev GA (2003b) The spiders of Serbia. Inst. Zoology, Fac. Biology, Belgrade & Inst. Zoology, Bulg. Acad. Sci., Sofia. 832 pp.
Deltshev C, Komnenov M, Blagoev G, Georgiev T, Lazarov S, Stojkoska E, Naumova M (2013) Faunistic diversity of spiders (Araneae) in Galichitsa Mountain (FYR Macedonia). Biodiversity Data Journal 1(e977): 1-70
Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp.
Grimm U (1985) Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF) 26: 1-318
Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204
Ivanov V V (2013) The checklist of Belarusian spiders (Arachnida, Araneae). Zoology and Ecology 4: 293-311
Izmailova M V (1989) [Fauna of Spiders of South Part of Eastern Siberia]. Irkutsk State University Publishing, 184 pp
Jézéquel J F (1962a) Contribution à l'étude des Zelotes femelles (Araneida, Labidognatha, Drassodidae "Gnaphosidae") de la fauna française. Verhandlungen der Deutschen Zoologischen Gesellschaft (Zool. Anz.) 25(Suppl.): 519-532
Koponen S, Fritzén N R, Pajunen T (2016) Checklist of spiders in Finland (Araneae). 6th version, December 2016, online at http://biolcoll.utu.fi/arach/checklist_of_spiders_in_Finland.htm
Kostanjšek R (unpubl.)
Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91
Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001]
Le Peru B (2007) Catalogue et répartition des araignées de France. Revue Arachnologique 16: 1-468
Løvbrekke H (unpubl.)
Maurer R, Hänggi A (1990) Katalog der schweizerischen Spinnen. Documenta Faunistica Helvetiae 12
Miller F (1967) Studien über die Kopulationsorgane der Spinnengattung Zelotes, Micaria, Robertus und Dipoena nebst Beschreibung einiger neuen oder unvollkommen bekannten Spinnenarten. Přírodovědné práce ústavů Československé Akademie Věd v Brně (N.S.) 1: 251-298
Milosevic B (2002) Aranea – Popis vrsta – Checklist. Hrvatsko entomološko društvo – Inventar entomofaune Hrvatske [Aranea – List of species – Checklist. Croatian Entomological Society – Inventory of Croatian entomofauna].
Morka G H (unpubl.)
Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App
Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html
Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html
Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184
Scharff N, Gudik-Sørensen O (2006) Katalog over Danmarks edderkopper (Araneae). Catalogue of the spiders of Denmark (Araneae). Entomologiske Meddelelser 74: 3-71
Thaler K (1989) Das Männchen von Zelotes devotus Grimm (Arachnida: Araneae, Gnaphosidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 62: 363-366
Thaler K, Knoflach B (2004b) Zur Faunistik der Spinnen (Araneae) von Österreich: Gnaphosidae, Thomisidae (Dionycha pro parte). Linzer Biologische Beiträge 36: 417-484
Tullgren A (1946) Svensk spindelfauna. 3. Egentiliga spinddlar. Araneae Fam 5-7. Clubionidae, Zoridae och Gnaphosidae. Stockholm (Entomol Fören.): 141 pp.
Tuneva T K, Esyunin S L (2003a) A review of the Gnaphosidae fauna of the Urals (Aranei), 3. New species and new records, chiefly from the south Urals. Arthropoda Selecta 11: 223-234
Vilbaste A (1987) Eesti Ämblikud (Aranei). Annoteeritud nimestik. Estonian spiders (Aranei). An annotated check list. Valgus, Tallinn. 113 pp. & 512 maps on 67 unpag. pages
Weiss I, Petrișor A (1999) List of the spiders (Arachnida: Araneae) from Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 41: 79-107
WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (13.2.2026) doi: 10.24436/2
Updates
| 05-01-2026 | Image insert | |
| 10-08-2025 | Distribution insert | Detail |
| 10-08-2025 | Distribution update | Detail |
| 20-12-2023 | Distribution update | Detail |
| 27-09-2022 | Image insert | |
| 08-03-2022 | Image insert | |
| 26-08-2020 | Distribution update | Detail |
| 02-04-2017 | Image insert | |
| 16-08-2016 | Image insert | |
| 02-02-2016 | Image insert | |
| 05-02-2015 | Image insert | |
| 15-10-2013 | Distribution update | Detail |