Gnaphosidae  Pocock, 1898 / Haplodrassus  Chamberlin, 1922   43 species

Haplodrassus umbratilis  (L. Koch, 1866)

Description

Male palp with plate-like bulge of distal bulbal apophysis, situated basally. Tibial apophysis somewhat widened distally. Epigyne with slantingly forward-directed protrusion. Prosoma light brown. Chelicerae darker than prosoma. Distal leg segments darker than prosoma. Opisthosoma light brown.

Body length male: 3.9-6 mm
Body length female: 5.1-7.5 mm
Additional information

in dry and open pine forests and heathland, on the ground and under stones during the day

Frequency: Widely distributed

Distribution
  Presence
  No data
Global distribution (WSC 2020): Europe to Kazakhstan
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Crimea: ♂♀ – V-VII, ♂♂ – IV, ♀♀ – VIII, X
Figures
Distribution List
Austria   (no references) ||| Belarus   (Ivanov, 2013) ||| Belgium   (no references) ||| Bulgaria   (Blagoev et al., 2018) ||| Czechia   (no references) ||| Estonia   (no references) ||| Finland   (Koponen et al., 2016) ||| France   (no references) ||| Germany   (no references) ||| Greece   (no references) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Pantini & Isaia, 2019) ||| Latvia   (Cera, 2018) ||| Liechtenstein   (no references) ||| Lithuania   (no references) ||| Luxembourg   (Hermann, 1998; Kreuels et al., 2019) ||| Montenegro   (no references) ||| Netherlands   (no references) ||| North Macedonia   (no references) ||| Norway   (Aakra & Hauge, 2003) ||| Poland   (no references) ||| Portugal   (Branco et al., 2019) ||| Romania   (no references) ||| Russia, Central   (no references) ||| Russia, Eastern   (no references) ||| Russia, Northern   (no references) ||| Russia, Southern   (Otto, 2020) ||| Serbia   (no references) ||| Slovakia   (no references) ||| Spain   (Branco et al., 2019) ||| Sweden   (Kronestedt, 2001) ||| Switzerland   (no references) ||| Turkey (Asia)   (Danışman et al., 2019) ||| Ukraine   (Polchaninova & Prokopenko, 2019) ||| United Kingdom   (no references) |||
References

Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129 pdf

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). [N.B.: few records from this paper are not taken over and not commented] Zootaxa 4614: 201-254 pdf

Brændegaard J (1966) Edderkopper: Eller Spindlere I Danmarks Fauna 72: 1-224 pdf

Cera I (2018) The checklist of Latvian spiders (Arachnida: Araneae). Environmental and Experimental Biology 16: 139-152 pdf

Danışman T, Kunt K B, Özkütük R S (2019) The checklist of the spiders of Turkey. Version 2019 [last updated 1 June 2019], online at http://www.spidersofturkey.info pdf

Fjellberg A (unpubl.)

Hermann E (1998) Die Spinnen (Araneae) ausgewählter Halbtrockenrasen im Osten Luxemburgs. Bulletin de la Société des naturalistes Luxembourgeois 99: 189-199 pdf

Ivanov V V (2013) The checklist of Belarusian spiders (Arachnida, Araneae). Zoology and Ecology 4: 293-311 pdf

Koponen S, Fritzén N R, Pajunen T (2016) Checklist of spiders in Finland (Araneae). 6th version, December 2016, online at http://biolcoll.utu.fi/arach/checklist_of_spiders_in_Finland.htm pdf

Kovblyuk M M, Kastrygina Z A, Omelko M M (2012b) A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae,  Gnaphosidae). Zookeys 205: 59-89 pdf

Kreuels M, Staudt A, Christian S (2019) Die Spinnenfauna von Luxemburg – eine Zusammenstellung der Nachweise aus den Jahren 1906-2018 (Arachnida: Araneae). Bulletin de la Société des naturalistes luxembourgeois 121: 203-230 pdf

Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001] pdf

Locket G H, Millidge A F, Merrett P (1974) British Spiders, Volume III. Ray Society, London, 315 pp. pdf

Miller F (1971) Pavouci-Araneida. Klíč zvířeny ČSSR 4: 51-306 pdf

Miller F, Buchar J (1977) Neue Spinnenarten aus den Gattungen Zelotes Distel und Haplodrassus Chamberlin (Araneae, Gnaphosidae). Acta Universitatis Carolinae Biologica 1974: 157-171 pdf

Otto S (2020) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 03.2020. Internet: caucasus-spiders.info.

Pantini P, Isaia M (2019) Araneae.it: the online catalog of Italian spiders, with addenda on other arachnid orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). http://araneae.it. Fragmenta Entomologica 51: 127-152 pdf

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V, Shmatko V Y (2017) A new species of spiders of the genus Haplodrassus Chamberlin, 1922 (Aranei: Gnaphosidae) from the northern pre-Caspian area. Caucasian Entomological Bulletin 13: 11-14 pdf

Roberts M J (1995) Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp. pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Tullgren A (1946) Svensk spindelfauna. 3. Egentiliga spinddlar. Araneae Fam 5-7. Clubionidae, Zoridae och Gnaphosidae. Stockholm (Entomol Fören.): 141 pp. pdf

World Spider Catalog (2020) World Spider Catalog. Version 21.0. Natural History Museum Bern, online at http://wsc.nmbe.ch (20.03.2020) doi: 10.24436/2 pdf

Updates
17-03-2020 Distribution insert Detail
01-01-2020 Image insert
02-10-2017 Image insert