Gnaphosidae  Pocock, 1898 / Haplodrassus  Chamberlin, 1922   43 species

Haplodrassus signifer  (C. L. Koch, 1839)

Description

Male palp with distinctly keeled distal bulbal apophysis. Prosoma red-brown, eye region darker. Chelicerae darker than prosoma. Legs brighter than prosoma. Opisthosoma provided with copper-coloured hairs.

Body length male: 4.7-7.4 mm
Body length female: 5.1-11.8 mm
Additional information

On dry meadows, on sand and stony debris, lawns, heathland, open forests and damp areas, up to 3000 m in the Alps, in litter and under stones during the day

Frequency: Very frequent, widely distributed

Distribution
  Presence
  No data
Global distribution (WSC 2020): North America, Europe, North Africa, Turkey, Caucasus, Russia (Europe to Far East), Israel, Iran, Central Asia, China, Korea
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List
Albania   (Deltshev et al., 2011b) ||| Algeria   (Bosmans, unpubl.) ||| Austria   (no references) ||| Azerbaijan   (Otto, 2020) ||| Belarus   (Ivanov, 2013) ||| Belgium   (no references) ||| Bulgaria   (Blagoev et al., 2018) ||| Croatia   (no references) ||| Cyprus   (Bosmans et al., 2019c) ||| Czechia   (no references) ||| Denmark   (no references) ||| Estonia   (no references) ||| Faroe Islands   (Lissner et al., 2016b) ||| Finland   (Koponen et al., 2016) ||| France   (no references) ||| France / Corsica   (no references) ||| Georgia   (Otto, 2020) ||| Germany   (no references) ||| Greece   (no references) ||| Greece / Crete   (no references) ||| Hungary   (Samu & Szinetár, 1999) ||| Iceland   (Agnarsson, 1996) ||| Ireland   (no references) ||| Italy   (Pantini & Isaia, 2019) ||| Italy / Sardinia   (Pantini & Isaia, 2019) ||| Italy / Sicily   (Pantini & Isaia, 2019) ||| Kosovo   (no references) ||| Latvia   (Cera, 2018) ||| Liechtenstein   (no references) ||| Lithuania   (no references) ||| Luxembourg   (Hermann, 1998; Kreuels et al., 2019) ||| Malta   (Pfliegler et al., 2017) ||| Moldova   (no references) ||| Montenegro   (no references) ||| Morocco   (Bosmans, unpubl.) ||| Netherlands   (no references) ||| North Macedonia   (no references) ||| Northern Ireland   (no references) ||| Norway   (Aakra & Hauge, 2003) ||| Poland   (no references) ||| Portugal   (Branco et al., 2019) ||| Romania   (no references) ||| Russia, Central   (no references) ||| Russia, Eastern   (Tuneva & Esyunin, 2003a) ||| Russia, Kaliningrad Region   (no references) ||| Russia, Northern   (no references) ||| Russia, Southern   (Otto, 2020) ||| Russia, Western   (no references) ||| Serbia   (no references) ||| Slovakia   (no references) ||| Slovenia   (Kostanjšek & Kuntner, 2015) ||| Spain   (Branco et al., 2019) ||| Sweden   (Kronestedt, 2001) ||| Switzerland   (no references) ||| Tunisia   (Bosmans, unpubl.) ||| Turkey (Asia)   (Danışman et al., 2019) ||| Turkey (Europe)   (Danışman et al., 2019) ||| Ukraine   (Polchaninova & Prokopenko, 2019) ||| United Kingdom   (no references) |||
References

Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129 pdf

Agnarsson I (1996) Íslenskar köngulær Fjölrit Náttúrufræðistofnunar 31: 1-175 pdf

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Bosmans R, Kherbouche-Abrous O, Benhalima S, Hervé C (2018) The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the Maghreb in particular (Araneae: Gnaphosidae) Zootaxa 4451: 1-67 pdf

Bosmans R, Van Keer J, Russell-Smith A, Hadjiconstantis M, Komnenov M, Bosselaers J, Huber S, McCowan D, Snazell R, Decae A, Zoumides C, Kielhorn K-H, Oger P (2019c) Spiders of Cyprus (Araneae). A catalogue of all currently known species from Cyprus. Newsletter of the Belgian Arachnological Society 34, Suppl. 1: 1-173 pdf

Bosmans R. (unpubl.) Provisional list of spiders of North Africa. Database excerpt Aug. 2019

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). [N.B.: few records from this paper are not taken over and not commented] Zootaxa 4614: 201-254 pdf

Breitling R (2020) South European spiders from the Duffey collection in the Manchester Museum (Arachnida: Araneae). Arachnology 18: 333-362 pdf

Brændegaard J (1966) Edderkopper: Eller Spindlere I Danmarks Fauna 72: 1-224 pdf

Cera I (2018) The checklist of Latvian spiders (Arachnida: Araneae). Environmental and Experimental Biology 16: 139-152 pdf

Danışman T, Kunt K B, Özkütük R S (2019) The checklist of the spiders of Turkey. Version 2019 [last updated 1 June 2019], online at http://www.spidersofturkey.info pdf

Deltshev C, Vrenozi B, Blagoev G, Lazarov S (2011b) Spiders of Albania – faunistic and zoogeographical review (Arachnida: Araneae). Acta Zoologica Bulgarica 63: 125-144 pdf

Hermann E (1998) Die Spinnen (Araneae) ausgewählter Halbtrockenrasen im Osten Luxemburgs. Bulletin de la Société des naturalistes Luxembourgeois 99: 189-199 pdf

Ivanov V V (2013) The checklist of Belarusian spiders (Arachnida, Araneae). Zoology and Ecology 4: 293-311 pdf

Koponen S, Fritzén N R, Pajunen T (2016) Checklist of spiders in Finland (Araneae). 6th version, December 2016, online at http://biolcoll.utu.fi/arach/checklist_of_spiders_in_Finland.htm pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Kovblyuk M M, Kastrygina Z A, Omelko M M (2012b) A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae,  Gnaphosidae). Zookeys 205: 59-89 pdf

Kreuels M, Staudt A, Christian S (2019) Die Spinnenfauna von Luxemburg – eine Zusammenstellung der Nachweise aus den Jahren 1906-2018 (Arachnida: Araneae). Bulletin de la Société des naturalistes luxembourgeois 121: 203-230 pdf

Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001] pdf

Lissner J, Jensen J-K, Hansen L J, Simonsen W, Kelduni R, Nissen K (2016b) An updated checklist of spiders (Araneae) of the Faroe Islands. Norwegian Journal of Entomology 63: 197-240 pdf

Locket G H, Millidge A F (1951) British spiders. Vol. I. Ray Society, London, 310 pp. pdf

Løvbrekke H (unpubl.)

Noordam A P (1992) Gnaphosidae (Araneae) of the Netherlands, drawings, some notes, and some thoughts about ant-mimicry. viii & 16 pp. Published by the author. pdf

Otto S (2020) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 03.2020. Internet: caucasus-spiders.info.

Pantini P, Isaia M (2019) Araneae.it: the online catalog of Italian spiders, with addenda on other arachnid orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). http://araneae.it. Fragmenta Entomologica 51: 127-152 pdf

Pfliegler W A, Schönhofer A, Niedbała W, Vella P, Sciberras A, Vella A (2017) New records of mites (Acari) and harvestmen (Opiliones) from Malta with a preliminary checklist of Maltese Arachnida. Soil Organisms 89: 85-110 pdf

Platnick N I, Shadab M U (1975) A revision of the spider genera Haplodrassus and Orodrassus (Araneae, Gnaphosidae) in North America. American Museum Novitates 2583: 1-40 pdf

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Roberts M J (1995) Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp. pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Tullgren A (1946) Svensk spindelfauna. 3. Egentiliga spinddlar. Araneae Fam 5-7. Clubionidae, Zoridae och Gnaphosidae. Stockholm (Entomol Fören.): 141 pp. pdf

Tuneva T K, Esyunin S L (2003a) A review of the Gnaphosidae fauna of the Urals (Aranei), 3. New species and new records, chiefly from the south Urals. Arthropoda Selecta 11: 223-234 pdf

World Spider Catalog (2020) World Spider Catalog. Version 21.0. Natural History Museum Bern, online at http://wsc.nmbe.ch (20.03.2020) doi: 10.24436/2 pdf

Updates
26-08-2020 Distribution update Detail
04-05-2020 Image insert
17-03-2020 Distribution insert Detail
02-10-2018 Image insert
10-09-2018 Image insert
09-08-2018 Image insert
02-02-2016 Image insert
10-12-2013 Distribution update Detail
01-12-2013 Distribution update Detail