Gnaphosidae  Banks, 1892 / Scotophaeus  Simon, 1893   21 species

Scotophaeus scutulatus  (L. Koch, 1866)

Description

Tibial apophysis with distinct tip. Epigyne with ± square groove, distinctly longer than wide. Prosoma light to dark red-brown. Opisthosoma light to dark red-brown.

Body length male: 6.8-10.9 mm
Body length female: 8.3-15.9 mm
Additional information

mostly in and on buildings, rare in natural environment

Distribution
  Presence
  No data
Global distribution (WSC 2024): Europe, Algeria, Turkey, Caucasus, Russia (Europe to South Siberia), Iran, Central Asia
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List

"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.

Albania   (Deltshev et al., 2011b; Vrenozi, 2012; Blick, 2018) ||| Algeria   (Benhacene et al., 2023) ||| Austria   (no references) ||| Azerbaijan   (Otto, 2022) ||| Belgium   (Bosmans & Van Keer, 2017) ||| Bosnia and Herzegovina   (Komnenov, 2010) ||| Bulgaria   (Blagoev et al., 2018) ||| Croatia   (Milosevic, 2002) ||| Cyprus   (Bosmans et al., 2019c) ||| Czechia   (Růžička & Řezáč, 2022a) ||| Denmark   (no references) ||| France   (no references) ||| Georgia   (Otto, 2022) ||| Germany   (no references) ||| Greece   (no references) ||| Greece / Crete   (no references) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Pantini & Isaia, 2019) ||| Italy / Sardinia   (Pantini & Isaia, 2019) ||| Italy / Sicily   (Pantini & Isaia, 2019) ||| Liechtenstein   (Arnold, 2001) ||| Luxembourg   (Christian, 2015; Kreuels et al., 2019) ||| Malta   (Pfliegler et al., 2017) ||| Moldova   (Vasiliev & Mikhailov, 2023) ||| Netherlands   (no references) ||| North Macedonia   (Deltshev et al., 2013) ||| Poland   (Rozwałka & Stanska, 2008) ||| Portugal   (Branco et al., 2019) ||| Romania   (no references) ||| Russia, Central   (no references) ||| Russia, Southern   (Otto, 2022; Ponomarev, 2022) ||| Serbia   (no references) ||| Slovakia   (no references) ||| Slovenia   (Kostanjšek & Kuntner, 2015) ||| Spain   (Branco et al., 2019) ||| Sweden   (Kronestedt, 2001) ||| Switzerland   (no references) ||| Turkey (Asia)   (Danışman et al., 2024a) ||| Ukraine   (Polchaninova & Prokopenko, 2019) ||| United Kingdom   (Lavery, 2019) |||
References

Arnold K (2001) Beitrag zur Spinnenfauna (Arachnida, Araneae) des Fürstentums Liechtenstein. Berichte der Botanisch-Zoologischen Gesellschaft Liechtenstein-Sargans-Werdenberg 28: 211-244 pdf

Benhacene R, Adjami A, Hadjeb A, Kermiche K, Ouakid M L (2023) Bibliographic checklist of the Algerian spider fauna (Araneae). Zootaxa 5352: 301-357 pdf

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Blick T (2018) A small collection of spiders (Arachnida: Araneae) from the River Vjosa, Albania – with an updated spider checklist of Albania. Acta ZooBot Austria 155: 213-232 pdf

Bosmans R, Van Keer J, Russell-Smith A, Hadjiconstantis M, Komnenov M, Bosselaers J, Huber S, McCowan D, Snazell R, Decae A, Zoumides C, Kielhorn K-H, Oger P (2019c) Spiders of Cyprus (Araneae). A catalogue of all currently known species from Cyprus. Newsletter of the Belgian Arachnological Society 34, Suppl. 1: 1-173 pdf

Bosmans R, Van Keer K (2017) Een herziene soortenlijst van de Belgische spinnen (Araneae). Nieuwsbrief van de Belgische Arachnologische Vereniging 32: 39-69 pdf

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254 pdf

Brændegaard J (1966) Edderkopper: Eller Spindlere I Danmarks Fauna 72: 1-224 pdf

Chatzaki M, Thaler K, Mylonas M (2002b) Ground spiders (Gnaphosidae, Araneae) of Crete and adjacent areas of Greece. Taxonomy and distribution. II. Revue Suisse de Zoologie 109: 603-633 pdf

Christian S (2015) Micaria subopaca Westring, 1861, Scotophaeus blackwalli (Thorell, 1871) and S. scutulatus (L.Koch, 1866): three species of gnaphosids new to Luxembourg (Arachnida, Gnaophosidae). Bulletin de la Société des naturalistes luxembourgeois 117: 87-90 pdf

Danışman T, Kunt K B, Özkütük R S, Coşar İ (2024a) The checklist of the spiders of Turkey. Version 2024 [last updated 01 January 2024], online at http://www.spidersofturkey.info pdf

Deltshev C, Komnenov M, Blagoev G, Georgiev T, Lazarov S, Stojkoska E, Naumova M (2013) Faunistic diversity of spiders (Araneae) in Galichitsa Mountain (FYR Macedonia). Biodiversity Data Journal 1(e977): 1-70 pdf

Deltshev C, Vrenozi B, Blagoev G, Lazarov S (2011b) Spiders of Albania – faunistic and zoogeographical review (Arachnida: Araneae). Acta Zoologica Bulgarica 63: 125-144 pdf

Komnenov M (2010) Checklist of spiders (Araneae) of Bosnia and Herzegovina. Uzizaž i Biospeld 5(2009): 52-69 pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Kreuels M, Staudt A, Christian S (2019) Die Spinnenfauna von Luxemburg – eine Zusammenstellung der Nachweise aus den Jahren 1906-2018 (Arachnida: Araneae). Bulletin de la Société des naturalistes luxembourgeois 121: 203-230 pdf

Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001] pdf

Lavery A (2019) A revised checklist of the spiders of Great Britain and Ireland. Arachnology 18: 196-212 pdf

Lecigne S (2021a) A new species of Sintula (Linyphiidae), redescription of Brigittea innocens (Dictynidae) and eight spider species newly recorded for Turkey (Araneae). Arachnologische Mitteilungen 62: 11-34 pdf

Miller F (1971) Pavouci-Araneida. Klíč zvířeny ČSSR 4: 51-306 pdf

Milosevic B (2002) Aranea – Popis vrsta – Checklist. Hrvatsko entomološko društvo – Inventar entomofaune Hrvatske [Aranea – List of species – Checklist. Croatian Entomological Society – Inventory of Croatian entomofauna]. pdf

Noordam A P (1992) Gnaphosidae (Araneae) of the Netherlands, drawings, some notes, and some thoughts about ant-mimicry. viii & 16 pp. Published by the author. pdf

Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.

Pantini P, Isaia M (2019) Araneae.it: the online catalog of Italian spiders, with addenda on other arachnid orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). Fragmenta Entomologica 51: 127-152 pdf

Pfliegler W A, Schönhofer A, Niedbała W, Vella P, Sciberras A, Vella A (2017) New records of mites (Acari) and harvestmen (Opiliones) from Malta with a preliminary checklist of Maltese Arachnida. Soil Organisms 89: 85-110 pdf

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Roberts M J (1995) Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp. pdf

Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Tullgren A (1946) Svensk spindelfauna. 3. Egentiliga spinddlar. Araneae Fam 5-7. Clubionidae, Zoridae och Gnaphosidae. Stockholm (Entomol Fören.): 141 pp. pdf

Vasiliev A I, Mikhailov K G (2023) Annotated checklist of spiders (Araneae) of the Republic of Moldova. Invertebrate Zoology 20: 307-342 pdf

Vrenozi B (2012) A collection of spiders (Araneae) in Albanian coastal areas. Arachnologische Mitteilungen 44: 41-46 pdf

WSC (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, online at http://wsc.nmbe.ch (22.01.2024) doi: 10.24436/2 pdf

Updates
30-05-2023 Distribution update Detail
08-07-2021 Image insert
08-04-2021 Distribution insert Detail
15-07-2020 Distribution update Detail
08-06-2020 Distribution update Detail
20-05-2020 Distribution update Detail
15-10-2019 Distribution update Detail