Gnaphosidae  Pocock, 1898 / Scotophaeus  Simon, 1893   18 species

Scotophaeus scutulatus  (L. Koch, 1866)

Description

Tibial apophysis with distinct tip. Epigyne with ± square groove, distinctly longer than wide. Prosoma light to dark red-brown. Opisthosoma light to dark red-brown.

Body length male: 6.8-10.9 mm
Body length female: 8.3-15.9 mm
Additional information

mostly in and on buildings, rare in natural environment

Distribution
  Presence
  No data
Global distribution (WSC 2020): Europe, Algeria, Turkey, Caucasus, Russia (Europe to South Siberia), Iran, Central Asiaia), Central Asia
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List
Albania   (no references) ||| Algeria   (Bosmans, unpubl.) ||| Austria   (no references) ||| Azerbaijan   (Otto, 2020) ||| Belgium   (no references) ||| Bosnia and Herzegovina   (Komnenov, 2010) ||| Bulgaria   (Blagoev et al., 2018) ||| Croatia   (no references) ||| Cyprus   (Bosmans et al., 2019c) ||| Czechia   (no references) ||| Denmark   (no references) ||| France   (no references) ||| Georgia   (Otto, 2020) ||| Germany   (no references) ||| Greece   (no references) ||| Greece / Crete   (no references) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Pantini & Isaia, 2019) ||| Italy / Sardinia   (Pantini & Isaia, 2019) ||| Liechtenstein   (no references) ||| Luxembourg   (Christian, 2015; Kreuels et al., 2019) ||| Malta   (Pfliegler et al., 2017) ||| Moldova   (no references) ||| Montenegro   (no references) ||| Netherlands   (no references) ||| North Macedonia   (no references) ||| Poland   (no references) ||| Portugal   (Branco et al., 2019) ||| Romania   (no references) ||| Russia, Central   (no references) ||| Russia, Southern   (Otto, 2020) ||| Serbia   (no references) ||| Slovakia   (no references) ||| Slovenia   (Kostanjšek & Kuntner, 2015) ||| Spain   (Branco et al., 2019) ||| Sweden   (Kronestedt, 2001) ||| Switzerland   (no references) ||| Turkey (Asia)   (Danışman et al., 2019) ||| Ukraine   (Polchaninova & Prokopenko, 2019) ||| United Kingdom   (no references) |||
References

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Bosmans R, Van Keer J, Russell-Smith A, Hadjiconstantis M, Komnenov M, Bosselaers J, Huber S, McCowan D, Snazell R, Decae A, Zoumides C, Kielhorn K-H, Oger P (2019c) Spiders of Cyprus (Araneae). A catalogue of all currently known species from Cyprus. Newsletter of the Belgian Arachnological Society 34, Suppl. 1: 1-173 pdf

Bosmans R. (unpubl.) Provisional list of spiders of North Africa. Database excerpt Aug. 2019

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). [N.B.: few records from this paper are not taken over and not commented] Zootaxa 4614: 201-254 pdf

Brændegaard J (1966) Edderkopper: Eller Spindlere I Danmarks Fauna 72: 1-224 pdf

Chatzaki M, Thaler K, Mylonas M (2002b) Ground spiders (Gnaphosidae, Araneae) of Crete and adjacent areas of Greece. Taxonomy and distribution. II. Revue Suisse de Zoologie 109: 603-633 pdf

Christian S (2015) Micaria subopaca Westring, 1861, Scotophaeus blackwalli (Thorell, 1871) and S. scutulatus (L.Koch, 1866): three species of gnaphosids new to Luxembourg (Arachnida, Gnaophosidae). Bulletin de la Société des naturalistes luxembourgeois 117: 87-90 pdf

Danışman T, Kunt K B, Özkütük R S (2019) The checklist of the spiders of Turkey. Version 2019 [last updated 1 June 2019], online at http://www.spidersofturkey.info pdf

Komnenov M (2010) Checklist of spiders (Araneae) of Bosnia and Herzegovina. Uzizaž i Biospeld 5(2009): 52-69 pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Kreuels M, Staudt A, Christian S (2019) Die Spinnenfauna von Luxemburg – eine Zusammenstellung der Nachweise aus den Jahren 1906-2018 (Arachnida: Araneae). Bulletin de la Société des naturalistes luxembourgeois 121: 203-230 pdf

Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001] pdf

Miller F (1971) Pavouci-Araneida. Klíč zvířeny ČSSR 4: 51-306 pdf

Noordam A P (1992) Gnaphosidae (Araneae) of the Netherlands, drawings, some notes, and some thoughts about ant-mimicry. viii & 16 pp. Published by the author. pdf

Otto S (2020) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 03.2020. Internet: caucasus-spiders.info.

Pantini P, Isaia M (2019) Araneae.it: the online catalog of Italian spiders, with addenda on other arachnid orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). http://araneae.it. Fragmenta Entomologica 51: 127-152 pdf

Pfliegler W A, Schönhofer A, Niedbała W, Vella P, Sciberras A, Vella A (2017) New records of mites (Acari) and harvestmen (Opiliones) from Malta with a preliminary checklist of Maltese Arachnida. Soil Organisms 89: 85-110 pdf

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Roberts M J (1995) Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp. pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Tullgren A (1946) Svensk spindelfauna. 3. Egentiliga spinddlar. Araneae Fam 5-7. Clubionidae, Zoridae och Gnaphosidae. Stockholm (Entomol Fören.): 141 pp. pdf

World Spider Catalog (2020) World Spider Catalog. Version 21.0. Natural History Museum Bern, online at http://wsc.nmbe.ch (20.03.2020) doi: 10.24436/2 pdf

Updates
15-07-2020 Distribution update Detail
08-06-2020 Distribution update Detail
20-05-2020 Distribution update Detail
15-10-2019 Distribution update Detail