Gnaphosidae  Banks, 1892 / Scotophaeus  Simon, 1893   21 species

Scotophaeus scutulatus (L. Koch, 1866)

Description

Tibial apophysis with distinct tip. Epigyne with ± square groove, distinctly longer than wide. Prosoma light to dark red-brown. Opisthosoma light to dark red-brown.

Body length male: 6.8-10.9 mm
Body length female: 8.3-15.9 mm
Additional information

mostly in and on buildings, rare in natural environment

Distribution
  Presence
  No data
Global distribution (WSC 2026): Europe, Algeria, Turkey, Caucasus, Russia (Europe to South Siberia), Iran, Central Asia
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List

"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.

Albania   (Deltshev et al., 2011b; Vrenozi, 2012; Blick, 2018) ||| Algeria   (Benhacene et al., 2023) ||| Austria   (Thaler & Knoflach, 2004b) ||| Azerbaijan   (Otto, 2022) ||| Belgium   (Bosmans & Van Keer, 2017) ||| Bosnia and Herzegovina   (Komnenov, 2010) ||| Bulgaria   (Blagoev et al., 2018) ||| Croatia   (Milosevic, 2002) ||| Cyprus   (Bosmans et al., 2019c) ||| Czechia   (Růžička & Řezáč, 2022a) ||| Denmark   (Scharff & Gudik-Sørensen, 2006) ||| France   (Le Peru, 2007) ||| Georgia   (Otto, 2022) ||| Germany   (Blick et al., 2016) ||| Greece   (Bosmans & Chatzaki, 2005) ||| Greece / Crete   (Bosmans & Chatzaki, 2005) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Isaia et al., 2025) ||| Italy / Sardinia   (Isaia et al., 2025) ||| Italy / Sicily   (Isaia et al., 2025) ||| Liechtenstein   (Arnold, 2001) ||| Lithuania   (Biteniekytė & Rėlys, 2012) ||| Luxembourg   (Christian, 2015; Kreuels et al., 2019) ||| Malta   (Pfliegler et al., 2017) ||| Moldova   (Vasiliev & Mikhailov, 2023) ||| Netherlands   (van Helsdingen, 2025a) ||| North Macedonia   (Deltshev et al., 2013) ||| Poland   (Rozwałka & Stanska, 2008) ||| Portugal   (Branco et al., 2019) ||| Romania   (Weiss & Petrișor, 1999) ||| Russia, Central   (no references) ||| Russia, Southern   (Otto, 2022; Ponomarev, 2022) ||| Serbia   (Stanković, 2023) ||| Slovakia   (Gajdoš et al., 1999) ||| Slovenia   (Kostanjšek & Kuntner, 2015) ||| Spain   (Branco et al., 2019) ||| Sweden   (Kronestedt, 2001) ||| Switzerland   (Maurer & Hänggi, 1990) ||| Turkey (Asia)   (Danışman et al., 2025a) ||| Ukraine   (Polchaninova & Prokopenko, 2019) ||| United Kingdom   (Lavery, 2019) |||
References

Almquist S (2007) Swedish Araneae, part 2 – families Dictynidae to Salticidae. Insect Systematics & Evolution, Supplement 63(2006): 285-601 pdf

Arnold K (2001) Beitrag zur Spinnenfauna (Arachnida, Araneae) des Fürstentums Liechtenstein. Berichte der Botanisch-Zoologischen Gesellschaft Liechtenstein-Sargans-Werdenberg 28: 211-244 pdf

Benhacene R, Adjami A, Hadjeb A, Kermiche K, Ouakid M L (2023) Bibliographic checklist of the Algerian spider fauna (Araneae). Zootaxa 5352: 301-357 pdf

Biteniekytė M, Rėlys V (2012) The checklist of Lithuanian spiders (Arachnida: Araneae). Biologija 57: 148-158 pdf

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Blick T (2018) A small collection of spiders (Arachnida: Araneae) from the River Vjosa, Albania – with an updated spider checklist of Albania. Acta ZooBot Austria 155: 213-232 pdf

Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510 pdf

Bosmans R, Chatzaki M (2005) A catalogue of the spiders of Greece – a critical review of all spider species cited from Greece with their localities. Nieuwsbrief van de Belgische arachnologische Vereniging 20 (2, suppl.): 1-124 pdf

Bosmans R, Van Keer J, Russell-Smith A, Hadjiconstantis M, Komnenov M, Bosselaers J, Huber S, McCowan D, Snazell R, Decae A, Zoumides C, Kielhorn K-H, Oger P (2019c) Spiders of Cyprus (Araneae). A catalogue of all currently known species from Cyprus. Newsletter of the Belgian Arachnological Society 34, Suppl. 1: 1-173 pdf

Bosmans R, Van Keer K (2017) Een herziene soortenlijst van de Belgische spinnen (Araneae). Nieuwsbrief van de Belgische Arachnologische Vereniging 32: 39-69 pdf

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254 pdf

Brændegaard J (1966) Edderkopper: Eller Spindlere I Danmarks Fauna 72: 1-224 pdf

Chatzaki M, Thaler K, Mylonas M (2002b) Ground spiders (Gnaphosidae, Araneae) of Crete and adjacent areas of Greece. Taxonomy and distribution. II. Revue Suisse de Zoologie 109: 603-633 pdf

Christian S (2015) Micaria subopaca Westring, 1861, Scotophaeus blackwalli (Thorell, 1871) and S. scutulatus (L.Koch, 1866): three species of gnaphosids new to Luxembourg (Arachnida, Gnaophosidae). Bulletin de la Société des naturalistes luxembourgeois 117: 87-90 pdf

Danışman T, Kunt K B, Özkütük R S, Coşar İ (2025a) The checklist of the spiders of Turkey. Version 2025 [last updated 22 July 2025], online at http://www.spidersofturkey.info pdf

Deltshev C, Komnenov M, Blagoev G, Georgiev T, Lazarov S, Stojkoska E, Naumova M (2013) Faunistic diversity of spiders (Araneae) in Galichitsa Mountain (FYR Macedonia). Biodiversity Data Journal 1(e977): 1-70 pdf

Deltshev C, Vrenozi B, Blagoev G, Lazarov S (2011b) Spiders of Albania – faunistic and zoogeographical review (Arachnida: Araneae). Acta Zoologica Bulgarica 63: 125-144 pdf

Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp.

Grimm U (1985) Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF) 26: 1-318 pdf

Helsdingen P J van (2025a) Catalogus van de Nederlandse spinnen. Versie 2025.1. 249 pp. pdf

Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204 pdf

Komnenov M (2010) Checklist of spiders (Araneae) of Bosnia and Herzegovina. Uzizaž i Biospeld 5(2009): 52-69 pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Kreuels M, Staudt A, Christian S (2019) Die Spinnenfauna von Luxemburg – eine Zusammenstellung der Nachweise aus den Jahren 1906-2018 (Arachnida: Araneae). Bulletin de la Société des naturalistes luxembourgeois 121: 203-230 pdf

Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001] pdf

Lavery A (2019) A revised checklist of the spiders of Great Britain and Ireland. Arachnology 18: 196-212 pdf

Le Peru B (2007) Catalogue et répartition des araignées de France. Revue Arachnologique 16: 1-468 pdf

Lecigne S (2021a) A new species of Sintula (Linyphiidae), redescription of Brigittea innocens (Dictynidae) and eight spider species newly recorded for Turkey (Araneae). Arachnologische Mitteilungen 62: 11-34 pdf

Maurer R, Hänggi A (1990) Katalog der schweizerischen Spinnen. Documenta Faunistica Helvetiae 12 pdf

Miller F (1971) Pavouci-Araneida. Klíč zvířeny ČSSR 4: 51-306 pdf

Milosevic B (2002) Aranea – Popis vrsta – Checklist. Hrvatsko entomološko društvo – Inventar entomofaune Hrvatske [Aranea – List of species – Checklist. Croatian Entomological Society – Inventory of Croatian entomofauna]. pdf

Noordam A P (1992) Gnaphosidae (Araneae) of the Netherlands, drawings, some notes, and some thoughts about ant-mimicry. viii & 16 pp. Published by the author. pdf

Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.

Pfliegler W A, Schönhofer A, Niedbała W, Vella P, Sciberras A, Vella A (2017) New records of mites (Acari) and harvestmen (Opiliones) from Malta with a preliminary checklist of Maltese Arachnida. Soil Organisms 89: 85-110 pdf

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Roberts M J (1995) Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp. pdf

Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Scharff N, Gudik-Sørensen O (2006) Katalog over Danmarks edderkopper (Araneae). Catalogue of the spiders of Denmark (Araneae). Entomologiske Meddelelser 74: 3-71 pdf

Stanković B (2023) Diversity of spider fauna in the area of Jagodina (Serbia). In: Seydoşoğlu, S. & Durna Aydin, Ö (eds.) International Cukurova Agriculture and Veterinary Congress 4. Adana, pp. 590-615 pdf

Thaler K, Knoflach B (2004b) Zur Faunistik der Spinnen (Araneae) von Österreich: Gnaphosidae, Thomisidae (Dionycha pro parte). Linzer Biologische Beiträge 36: 417-484 pdf

Tullgren A (1946) Svensk spindelfauna. 3. Egentiliga spinddlar. Araneae Fam 5-7. Clubionidae, Zoridae och Gnaphosidae. Stockholm (Entomol Fören.): 141 pp. pdf

Vasiliev A I, Mikhailov K G (2023) Annotated checklist of spiders (Araneae) of the Republic of Moldova. Invertebrate Zoology 20: 307-342 pdf

Vrenozi B (2012) A collection of spiders (Araneae) in Albanian coastal areas. Arachnologische Mitteilungen 44: 41-46 pdf

Weiss I, Petrișor A (1999) List of the spiders (Arachnida: Araneae) from Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 41: 79-107 pdf

WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (13.2.2026) doi: 10.24436/2 pdf

Updates
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10-08-2025 Distribution insert Detail
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