Gnaphosidae  Pocock, 1898 / Scotophaeus  Simon, 1893   18 species

Scotophaeus scutulatus  (L. Koch, 1866)

Description

Tibial apophysis with distinct tip. Epigyne with ± square groove, distinctly longer than wide. Prosoma light to dark red-brown. Opisthosoma light to dark red-brown.

Body length male: 6.8-10.9 mm
Body length female: 8.3-15.9 mm
Additional information

mostly in and on buildings, rare in natural environment

Distribution
  Presence
  No data
Global distribution (WSC 2020): Europe, Algeria, Turkey, Caucasus, Russia (Europe to South Siberia), Iran, Central Asiaia), Central Asia
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List
Albania   ( no references ) ||| Algeria   ( Bosmans, unpubl.; ) ||| Austria   ( no references ) ||| Azerbaijan   ( Otto, 2020; ) ||| Belgium   ( no references ) ||| Bosnia and Herzegovina   ( no references ) ||| Bulgaria   ( Blagoev et al., 2018; ) ||| Croatia   ( no references ) ||| Cyprus   ( Bosmans et al., 2019c; ) ||| Czechia   ( no references ) ||| Denmark   ( no references ) ||| France   ( no references ) ||| Georgia   ( Otto, 2020; ) ||| Germany   ( no references ) ||| Greece   ( no references ) ||| Greece / Crete   ( no references ) ||| Hungary   ( no references ) ||| Italy   ( Pantini & Isaia, 2019; ) ||| Italy / Sardinia   ( Pantini & Isaia, 2019; ) ||| Liechtenstein   ( no references ) ||| Luxembourg   ( Christian, 2015; Kreuels et al., 2019; ) ||| Malta   ( Pfliegler et al., 2017; ) ||| Moldova   ( no references ) ||| Montenegro   ( no references ) ||| Netherlands   ( no references ) ||| North Macedonia   ( no references ) ||| Poland   ( no references ) ||| Portugal   ( no references ) ||| Romania   ( no references ) ||| Russia, Central   ( no references ) ||| Russia, Southern   ( Otto, 2020; ) ||| Serbia   ( no references ) ||| Slovakia   ( no references ) ||| Slovenia   ( Kostanjšek & Kuntner, 2015; ) ||| Spain   ( no references ) ||| Sweden   ( no references ) ||| Switzerland   ( no references ) ||| Turkey (Asia)   ( Danışman et al., 2019; ) ||| Ukraine   ( Polchaninova & Prokopenko, 2019; ) ||| United Kingdom   ( no references ) |||
References

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018)

Bosmans R, Van Keer J, Russell-Smith A, Hadjiconstantis M, Komnenov M, Bosselaers J, Huber S, McCowan D, Snazell R, Decae A, Zoumides C, Kielhorn K-H, Oger P (2019c) Spiders of Cyprus (Araneae). A catalogue of all currently known species from Cyprus. Newsletter of the Belgian Arachnological Society 34, Suppl. 1: 1-173 pdf

Bosmans R. (unpubl.) Provisional list of spiders of North Africa. Database excerpt Aug. 2019

Brændegaard J (1966) Edderkopper: Eller Spindlere I Danmarks Fauna 72: 1-224 pdf

Chatzaki M, Thaler K, Mylonas M (2002b) Ground spiders (Gnaphosidae, Araneae) of Crete and adjacent areas of Greece. Taxonomy and distribution. II. Rev Suisse Zool 109: 603-633

Christian S (2015) Micaria subopaca Westring, 1861, Scotophaeus blackwalli (Thorell, 1871) and S. scutulatus (L.Koch, 1866): three species of gnaphosids new to Luxembourg (Arachnida, Gnaophosidae). Bulletin de la Société des naturalistes luxembourgeois 117: 87-90 pdf

Danışman T, Kunt K B, Özkütük R S (2019) The checklist of the spiders of Turkey. Version 2019 [last updated 1 June 2019], online at http://www.spidersofturkey.info

Helsdingen P J van (2010a) Araneae. In: Fauna Europaea Database (Version 2010.1), online at www.european-arachnology.org

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Kreuels M, Staudt A, Christian S (2019) Die Spinnenfauna von Luxemburg – eine Zusammenstellung der Nachweise aus den Jahren 1906-2018 (Arachnida: Araneae). Bulletin de la Société des naturalistes luxembourgeois 121: 203-230 pdf

Miller F (1971) Pavouci-Araneida In: Daniel M & Cerny V. (Hrsg.) Klic zvireny CSSR, Dil. IV. Ceskoslovenska Akademie Ved, Prag: 51-306

Noordam A P (1992) Gnaphosidae (Araneae) of the Netherlands, drawings, some notes, and some thoughts about ant-mimicry. Published by the author.

Otto S (2020) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 03.2020. Internet: caucasus-spiders.info.

Pantini P, Isaia M (2019) Araneae.it: the online Catalog of Italian spiders, with addenda on other Arachnid Orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). http://araneae.it. Fragmenta Entomologica 51: 127-152 pdf

Pfliegler W A, Schönhofer A, Niedbała W, Vella P, Sciberras A, Vella A (2017) New records of mites (Acari) and harvestmen (Opiliones) from Malta with a preliminary checklist of Maltese Arachnida. Soil Organisms 89: 85-110 pdf

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Roberts M J (1995) Spiders of Britain and Northern Europe. Collins Field Guide Bath

Tullgren A (1946) Svensk spindelfauna. 3. Egentiliga spinddlar. Araneae Fam 5-7. Clubionidae, Zoridae och Gnaphosidae. Stockholm (Entomol Fören): 141 pp. pdf

World Spider Catalog (2020) World Spider Catalog. Version 21.0. Natural History Museum Bern, online at http://wsc.nmbe.ch (20.03.2020) doi: 10.24436/2

Updates
20-05-2020 Distribution update Detail
15-10-2019 Distribution update Detail