Gnaphosidae  Banks, 1892 / Drassyllus  Chamberlin, 1922   18 species

Drassyllus pusillus  (C. L. Koch, 1833)

Description
Male

Male palp with distal median apophysis, distally widened. Prosoma red-brown to black-brown. Metatarsus I-II light yellow. Tarsus light yellow. Opisthosoma red-brown to black-brown, with dark brown scuticula.

Body length male: 3.5-4.6 mm
Female

Epigyne with longitudinal cuticular ridge, distinctly sclerotised.

Body length female: 4.0-5.5 mm
Additional information

Found in very different habitats (forests, meadows, river-floodplains, heathland, sand and stony debris), under stones, in moss and leaf litter during the day

Distribution
  Presence
  No data
Global distribution (WSC 2024): Europe, Turkey, Caucasus, Russia (Europe to Far East), Iran, Central Asia, China
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List

"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.

Albania   (Naumova et al., 2016b) ||| Austria   (no references) ||| Azerbaijan   (Otto, 2022) ||| Belarus   (Ivanov, 2013) ||| Belgium   (Bosmans & Van Keer, 2017) ||| Bulgaria   (Blagoev et al., 2018) ||| Croatia   (Milosevic, 2002) ||| Cyprus   (Bosmans et al., 2019c) ||| Czechia   (Růžička & Řezáč, 2022a) ||| Denmark   (no references) ||| Estonia   (Vilbaste, 1987) ||| Finland   (Koponen et al., 2016) ||| France   (no references) ||| Georgia   (Otto, 2022) ||| Germany   (no references) ||| Greece   (no references) ||| Hungary   (Samu & Szinetár, 1999) ||| Ireland   (Lavery, 2019) ||| Italy   (Pantini & Isaia, 2019) ||| Latvia   (Cera, 2018) ||| Liechtenstein   (Wiedemeier, 1990) ||| Lithuania   (no references) ||| Luxembourg   (Hermann, 1998; Kreuels et al., 2019) ||| Malta   (Pfliegler et al., 2017) ||| Moldova   (Vasiliev & Mikhailov, 2023) ||| Netherlands   (no references) ||| North Macedonia   (Komnenov, 2014) ||| Northern Ireland   (Lavery, 2019) ||| Norway   (Aakra & Hauge, 2003) ||| Poland   (Rozwałka & Stanska, 2008) ||| Romania   (no references) ||| Russia, Central   (Ponomarev, 2022) ||| Russia, Eastern   (Tuneva & Esyunin, 2003a) ||| Russia, Kaliningrad Region   (no references) ||| Russia, Northern   (no references) ||| Russia, Southern   (Otto, 2022; Ponomarev, 2022) ||| Russia, Western   (no references) ||| Serbia   (no references) ||| Slovakia   (no references) ||| Slovenia   (Kostanjšek & Kuntner, 2015) ||| Spain   (Branco et al., 2019) ||| Sweden   (Kronestedt, 2001) ||| Switzerland   (no references) ||| Turkey (Asia)   (Danışman et al., 2024a) ||| Ukraine   (Polchaninova & Prokopenko, 2019) ||| United Kingdom   (Lavery, 2019) |||
References

Aakra K (unpubl.)

Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129 pdf

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Bosmans R, Van Keer J, Russell-Smith A, Hadjiconstantis M, Komnenov M, Bosselaers J, Huber S, McCowan D, Snazell R, Decae A, Zoumides C, Kielhorn K-H, Oger P (2019c) Spiders of Cyprus (Araneae). A catalogue of all currently known species from Cyprus. Newsletter of the Belgian Arachnological Society 34, Suppl. 1: 1-173 pdf

Bosmans R, Van Keer K (2017) Een herziene soortenlijst van de Belgische spinnen (Araneae). Nieuwsbrief van de Belgische Arachnologische Vereniging 32: 39-69 pdf

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254 pdf

Brændegaard J (1966) Edderkopper: Eller Spindlere I Danmarks Fauna 72: 1-224 pdf

Cera I (2018) The checklist of Latvian spiders (Arachnida: Araneae). Environmental and Experimental Biology 16: 139-152 pdf

Danışman T, Kunt K B, Özkütük R S, Coşar İ (2024a) The checklist of the spiders of Turkey. Version 2024 [last updated 01 January 2024], online at http://www.spidersofturkey.info pdf

Hermann E (1998) Die Spinnen (Araneae) ausgewählter Halbtrockenrasen im Osten Luxemburgs. Bulletin de la Société des naturalistes Luxembourgeois 99: 189-199 pdf

Ivanov V V (2013) The checklist of Belarusian spiders (Arachnida, Araneae). Zoology and Ecology 4: 293-311 pdf

Jézéquel J F (1962a) Contribution à l'étude des Zelotes femelles (Araneida, Labidognatha, Drassodidae "Gnaphosidae") de la fauna française. Verhandlungen der Deutschen Zoologischen Gesellschaft (Zool. Anz.) 25(Suppl.): 519-532 pdf

Komnenov M (2014) Spider fauna of the Osogovo Mt. Range, Northeastern Macedonia. Fauna Balkana 2: 1-267 pdf

Koponen S, Fritzén N R, Pajunen T (2016) Checklist of spiders in Finland (Araneae). 6th version, December 2016, online at http://biolcoll.utu.fi/arach/checklist_of_spiders_in_Finland.htm pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Kovblyuk M M (2003b) The spider genus Drassyllus Chamberlin, 1922 in the Crimean fauna, with description of a new species (Aranei: Gnaphosidae). Arthropoda Selecta 12: 23-28 pdf

Kreuels M, Staudt A, Christian S (2019) Die Spinnenfauna von Luxemburg – eine Zusammenstellung der Nachweise aus den Jahren 1906-2018 (Arachnida: Araneae). Bulletin de la Société des naturalistes luxembourgeois 121: 203-230 pdf

Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001] pdf

Lavery A (2019) A revised checklist of the spiders of Great Britain and Ireland. Arachnology 18: 196-212 pdf

Locket G H, Millidge A F (1951) British spiders. Vol. I. Ray Society, London, 310 pp. pdf

Løvbrekke H (unpubl.)

Miller F (1967) Studien über die Kopulationsorgane der Spinnengattung Zelotes, Micaria, Robertus und Dipoena nebst Beschreibung einiger neuen oder unvollkommen bekannten Spinnenarten. Přírodovědné práce ústavů Československé Akademie Věd v Brně (N.S.) 1: 251-298 pdf

Milosevic B (2002) Aranea – Popis vrsta – Checklist. Hrvatsko entomološko društvo – Inventar entomofaune Hrvatske [Aranea – List of species – Checklist. Croatian Entomological Society – Inventory of Croatian entomofauna]. pdf

Naumova M, Hristovski S, Hristov G H (2016b) Spiders (Arachnida: Araneae) from Prespa National Park, Albania. Acta Zoologica Bulgarica 68: 503-511 pdf

Noordam A P (1992) Gnaphosidae (Araneae) of the Netherlands, drawings, some notes, and some thoughts about ant-mimicry. viii & 16 pp. Published by the author. pdf

Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.

Pantini P, Isaia M (2019) Araneae.it: the online catalog of Italian spiders, with addenda on other arachnid orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). Fragmenta Entomologica 51: 127-152 pdf

Pfliegler W A, Schönhofer A, Niedbała W, Vella P, Sciberras A, Vella A (2017) New records of mites (Acari) and harvestmen (Opiliones) from Malta with a preliminary checklist of Maltese Arachnida. Soil Organisms 89: 85-110 pdf

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Roberts M J (1995) Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp. pdf

Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Tullgren A (1946) Svensk spindelfauna. 3. Egentiliga spinddlar. Araneae Fam 5-7. Clubionidae, Zoridae och Gnaphosidae. Stockholm (Entomol Fören.): 141 pp. pdf

Tuneva T K, Esyunin S L (2003a) A review of the Gnaphosidae fauna of the Urals (Aranei), 3. New species and new records, chiefly from the south Urals. Arthropoda Selecta 11: 223-234 pdf

Vasiliev A I, Mikhailov K G (2023) Annotated checklist of spiders (Araneae) of the Republic of Moldova. Invertebrate Zoology 20: 307-342 pdf

Vilbaste A (1987) Eesti Ämblikud (Aranei). Annoteeritud nimestik. Estonian spiders (Aranei). An annotated check list. Valgus, Tallinn. 113 pp. & 512 maps on 67 unpag. pages pdf

Wiedemeier P (1990) Die Bodenspinnen des Ruggeller Rietes. Berichte der Botanisch-Zoologischen Gesellschaft Liechtenstein-Sargans-Werdenberg 18: 319-344 pdf

WSC (2024) World Spider Catalog. Version 25.0. Natural History Museum Bern, online at http://wsc.nmbe.ch (22.01.2024) doi: 10.24436/2 pdf

Updates
15-04-2023 Image insert
15-01-2023 Distribution update Detail
27-09-2020 Image insert
26-08-2020 Distribution update Detail
28-07-2020 Image insert
17-03-2020 Distribution insert Detail
11-05-2018 Distribution update Detail