Phrurolithidae  Banks, 1892 / Phrurolithus  C. L. Koch, 1839   11 species

Phrurolithus pullatus Kulczyński, 1897

Description

Tibial apophysis with symmetric tip. Bulbus distally with 2 apophyses, overtopping embolus. Epigyne with laterally situated copulatory openings. Prosoma brown. Opisthosoma brown, with bright spots on dark ground.

Body length male: 1.8-2 mm
Body length female: 2.1-2.6 mm
Additional information

Dry meadows, open forests

Distribution
  Presence
  No data
Distribution comment: Portugal: mentioned by Branco et al. (2019), but interpreted here as probable misidentification, due to the general distribution of the species
Global distribution (WSC 2026): Central to Eastern Europe, Turkey, Caucasus, Iran, Kazakhstan, Central Asia, China
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List
Albania   (van Helsdingen & IJland, 2015) ||| Austria   (Grimm, 1986) ||| Azerbaijan   (Otto, 2022) ||| Bulgaria   (Blagoev et al., 2018) ||| Czechia   (Růžička & Řezáč, 2022a) ||| Georgia   (Ponomarev & Komarov, 2015) ||| Germany   (Blick et al., 2016) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy  Removed (Isaia et al., 2025) ||| North Macedonia   (Deltshev et al., 2013) ||| Poland   (Rozwałka & Stanska, 2008) ||| Romania   (Weiss & Petrișor, 1999) ||| Russia, Central   (no references) ||| Russia, Eastern   (Esyunin & Efimik, 1996a) ||| Russia, Southern   (Otto, 2022; Ponomarev & Khnykin, 2013; Ponomarev, 2022) ||| Slovakia   (Gajdoš et al., 1999) ||| Turkey (Asia)   (Danışman et al., 2017) ||| Ukraine   (Polchaninova & Prokopenko, 2019) |||
References

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510 pdf

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254 pdf

Danışman T, Varol M İ, Coşar İ (2017) Two new records of little-known myrmecomorphic spiders in Turkey (Araneae: Phrurolithidae and Zodariidae). Serket 15: 150-153 pdf

Deltshev C, Komnenov M, Blagoev G, Georgiev T, Lazarov S, Stojkoska E, Naumova M (2013) Faunistic diversity of spiders (Araneae) in Galichitsa Mountain (FYR Macedonia). Biodiversity Data Journal 1(e977): 1-70 pdf

Esyunin S L, Efimik V E (1996a) Catalogue of the spiders (Arachnida, Aranei) of the Urals. KMK Scientific Press, Moscow, 228 pp. pdf

Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp.

Grimm U (1986) Die Clubionidae Mitteleuropas: Corinninae und Liocranidae (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF) 27: 1-91 pdf

Helsdingen P J van, IJland S (2015) A quick scan of the spider fauna in the surroundings of Përmet, Albania (Arachnida, Araneae) - preliminary report. Nieuwsbrief SPINED 35: 15-33 pdf

Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204 pdf

Miller F (1971) Pavouci-Araneida. Klíč zvířeny ČSSR 4: 51-306 pdf

Mu Y N, Wang L Y, Shi S T, Zhang F (2025) New taxonomic and faunistic data on guardstone spiders (Araneae, Phrurolithidae) from China. Zootaxa 5719: 413-422 pdf

Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Ponomarev A V, Khnykin A S (2013) Spiders (Aranei) of Volgograd sity and its environs. The South of Russia: Ecology, Development 2013(4): 109-136 pdf

Ponomarev A V, Komarov Y E (2015) Spiders (Aranei) of the Republic of South Ossetia. The South of Russia: Ecology, Development 10(1): 116-147 pdf

Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Sterghiu C (1985) Fam. Clubionidae. Fauna Republicii Socialiste România (Arachnida) 5 (4): 1-168 pdf

Weiss I, Petrișor A (1999) List of the spiders (Arachnida: Araneae) from Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 41: 79-107 pdf

Wiehle H (1967) Beiträge zur Kenntnis der deutschen Spinnenfauna V. Senckenbergiana Biologica 48: 1-36 pdf

WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (20.5.2026) doi: 10.24436/2 pdf

Zamani A, Marusik Y M (2020b) A survey of Phrurolithidae (Arachnida: Araneae) in southern Caucasus, Iran and Central Asia. Zootaxa 4758: 311-329 pdf

Updates
13-11-2025 Image insert
22-07-2025 Distribution update Detail
20-09-2023 Distribution insert Detail
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29-12-2014 Distribution update Detail