Linyphiidae  Blackwall, 1859 / Scotargus  Simon, 1913   2 species

Scotargus pilosus Simon, 1913

Description
Male

Embolus large, describing a semi-circle, tip curved. Prosoma and legs yellowish orange. Opisthosoma pale grey.

Body length male: 2.7-3.1 mm
Female

Epigyne with convoluted scapus and broad epigynal plate. Colouration as in males.

Body length female: 2.3-3.8 mm
Additional information

in submountainous forests, partly microcavernicolous and in caves

Character states from linyphiid key
Distribution
  Presence
  No data
Global distribution (WSC 2026): Europe, Morocco, Algeria, Caucasus, Russia (Europe to Far East), Iran, Kazakhstan, Central Asia, Nepal
Figures
Distribution List
Algeria   (Bosmans, 2006a; Benhacene et al., 2023) ||| Austria   (Thaler, 1995) ||| Azerbaijan   (Otto, 2022) ||| Bulgaria   (Blagoev et al., 2018) ||| Czechia   (Růžička & Řezáč, 2022a) ||| France   (Le Peru, 2007) ||| Georgia   (Seropian et al., 2026) ||| Germany   (Blick et al., 2016) ||| Greece   (Tanasevitch, 2011a) ||| Greece / Crete   (Bosmans et al., 2013) ||| Italy   (Isaia et al., 2025) ||| Liechtenstein   (Arnold, 2001) ||| Montenegro   (Naumova et al., 2019a) ||| Morocco   (Lecigne et al., 2023) ||| North Macedonia   (Komnenov, 2014) ||| Poland   (Rozwałka & Stanska, 2008) ||| Russia, Northern   (no references) ||| Russia, Southern   (Otto, 2022; Ponomarev, 2022) ||| Serbia   (Ćurčić et al., 2003) ||| Slovakia   (Gajdoš et al., 1999) ||| Slovenia   (Kostanjšek & Kuntner, 2015) ||| Spain   (Branco et al., 2019) ||| Switzerland   (Maurer & Hänggi, 1990) ||| Ukraine   (Gajdoš et al., 2014) |||
References

Arnold K (2001) Beitrag zur Spinnenfauna (Arachnida, Araneae) des Fürstentums Liechtenstein. Berichte der Botanisch-Zoologischen Gesellschaft Liechtenstein-Sargans-Werdenberg 28: 211-244 pdf

Benhacene R, Adjami A, Hadjeb A, Kermiche K, Ouakid M L (2023) Bibliographic checklist of the Algerian spider fauna (Araneae). Zootaxa 5352: 301-357 pdf

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510 pdf

Bosmans R (2006a) Contribution to the knowledge of the Linyphiidae of the Maghreb. Part XI. Miscellaneous linyphiid genera and additions (Araneae: Linyphiidae: Linyphiinae). Bulletin & Annales de la Société Entomologique de Belgique 141: 125-161 pdf

Bosmans R, Van Keer J, Russell-Smith A, Kronestedt T, Alderweireldt M, Bosselaers J, De Koninck H (2013) Spiders of Crete (Araneae). A catalogue of all currently known species from the Greek island of Crete. Nieuwsbrief van de Belgische Arachnologische Vereniging 28 (Suppl. 1): 1-147 pdf

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254 pdf

Ćurčić B P M, Deltshev C C, Blagoev G A, Ćurčić S B, Tomić V T (2003) On the diversity of some spiders (Aranea: Arachnida) from Serbia. Archives of Biological Sciences Belgrade 55: 23-31 pdf

Gajdoš P, Moscaliuc L A, Rozwałka R, Hirna A, Majkus Z, Gubányi A, Heltai M G, Svatoň J (2014) Red List of spiders (Araneae) of the Carpathian Mts. pp. 118-171. In: Kadlečik J (ed.): Carpathian Red List of forest habitats and species. Carpathian list ofinvasive alien species (Draft). The State Nature Conservancy of the Slovak Republic, Banská Bystrica. pdf

Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp.

Helsdingen P J van (1973) On the affinity of the genus Scotargus Simon (Araneae: Linyphiidae). Bulletin of the British Arachnological Society 2: 189-192 pdf

Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204 pdf

Komnenov M (2014) Spider fauna of the Osogovo Mt. Range, Northeastern Macedonia. Fauna Balkana 2: 1-267 pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Le Peru B (2007) Catalogue et répartition des araignées de France. Revue Arachnologique 16: 1-468 pdf

Lecigne S, Moutaouakil S, Lips J (2023) Contribution to the knowledge of the spider fauna of Morocco (Arachnida: Araneae) – first note – on new records of cave spiders. Arachnologische Mitteilungen 66: 44-71 pdf

Maurer R, Hänggi A (1990) Katalog der schweizerischen Spinnen. Documenta Faunistica Helvetiae 12 pdf

Moritz M (1973) Neue und seltene Spinnen (Araneae) und Weberknechte (Opiliones) aus der DDR. Deutsche Entomologische Zeitschrift (N.F.) 20: 173-220 pdf

Naumova M, Lazarov S, Deltshev C (2019a) Faunistic diversity of the spiders in Montenegro (Arachnida: Araneae). Ecologica Montenegrina 22: 50-89 pdf

Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Seropian A, Bulbulashvili N, Egidunashvili V (2026) Spiders in a growing city: new country and regional records of spiders (Araneae) from Tbilisi, Georgia. Caucasiana 5: 31-39 pdf

Tanasevitch A V (2011a) On linyphiid spiders from the eastern and central Mediterranean kept at the Muséum d'histoire naturelle, Geneva. Revue Suisse de Zoologie 118: 49-91 pdf

Thaler K (1995) Beiträge zur Spinnenfauna von Nordtirol - 5. Linyphiidae 1: Linyphiinae (sensu Wiehle) (Arachnida: Araneida). Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck 82: 153-190 pdf

Wiehle H (1956) Spinnentiere oder Arachnoidea, X. 28: Familie Linyphiidae-Baldachinspinnen. Die Tierwelt Deutschlands 44: 1-337 pdf

WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (13.2.2026) doi: 10.24436/2 pdf

Zamani A, Darvishnia H, Marusik Y M (2023b) New data on cave spiders (Arachnida: Araneae) of Iran, with new species and records. Zootaxa 5361: 345-366 pdf

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