Linyphiidae  Blackwall, 1859 / Pityohyphantes  Simon, 1929   1 species

Pityohyphantes phrygianus (C. L. Koch, 1836)

Description

Male palp with long patellar apophysis. Paracymbium small. Epigyne with large, spoon-shaped scapus. Prosoma grey-brown, with narrow black margin and Y-shaped black median stripe. Sternum yellow, with red-brown margin. Chelicerae brown, anterior claw furrow margin with 3, posterior with 2 teeth. Legs yellow-brown, relatively long, with dark rings. Femur I-IV with blackish spotted ventral side. Opisthosoma light grey, slightly reddish, with broad, laterally indented median stripe. Lateral border red-brown or black, ventrally with dark, white margined median band.

Body length male: 4-6 mm
Body length female: 5.0-6.7 mm
Additional information

in shadowed forests of low mountain range, sporadically also in the lowlands

Character states from linyphiid key
Distribution
  Presence
  No data
Global distribution (WSC 2026): Europe, Russia (Europe to Far East), Kazakhstan, Japan
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List
Albania   (Deltshev et al., 2011b; Vrenozi, 2012; Blick, 2018) ||| Austria   (Thaler, 1995) ||| Belarus   (Ivanov, 2013) ||| Belgium   (Bosmans & Van Keer, 2017) ||| Bulgaria   (Blagoev et al., 2018) ||| Czechia   (Růžička & Řezáč, 2022a) ||| Denmark   (Scharff & Gudik-Sørensen, 2006) ||| Estonia   (Vilbaste, 1987) ||| Finland   (Koponen et al., 2016) ||| France   (Le Peru, 2007) ||| Germany   (Blick et al., 2016) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Isaia et al., 2025) ||| Italy / Sardinia   (Isaia et al., 2025) ||| Kosovo   (Geci et al., 2023) ||| Latvia   (Cera, 2018) ||| Liechtenstein   (Arnold, 2001) ||| Lithuania   (Biteniekytė & Rėlys, 2012) ||| Luxembourg   (Kreuels et al., 2019) ||| Montenegro   (Naumova et al., 2019a) ||| Netherlands   (van Helsdingen, 2025a) ||| North Macedonia   (Blagoev, 2002) ||| Norway   (Aakra & Hauge, 2003) ||| Poland   (Rozwałka & Stanska, 2008) ||| Romania   (Weiss & Petrișor, 1999) ||| Russia, Central   (Ponomarev, 2022) ||| Russia, Eastern   (Esyunin & Efimik, 1996a) ||| Russia, Kaliningrad Region   (Schikora, 2014) ||| Russia, Northern   (no references) ||| Russia, Southern   (Ponomarev, 2022) ||| Russia, Western   (no references) ||| Slovakia   (Gajdoš et al., 1999) ||| Slovenia   (Kostanjšek & Kuntner, 2015) ||| Sweden   (Kronestedt, 2001) ||| Switzerland   (Maurer & Hänggi, 1990) ||| Ukraine   (Polchaninova & Prokopenko, 2019) ||| United Kingdom   (Lavery, 2019) |||
References

Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129 pdf

Arnold K (2001) Beitrag zur Spinnenfauna (Arachnida, Araneae) des Fürstentums Liechtenstein. Berichte der Botanisch-Zoologischen Gesellschaft Liechtenstein-Sargans-Werdenberg 28: 211-244 pdf

Biteniekytė M, Rėlys V (2012) The checklist of Lithuanian spiders (Arachnida: Araneae). Biologija 57: 148-158 pdf

Blagoev G (2002) Check list of Macedonian spiders (Araneae). Acta Zoologica Bulgarica 54: 9-34 pdf

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Blick T (2018) A small collection of spiders (Arachnida: Araneae) from the River Vjosa, Albania – with an updated spider checklist of Albania. Acta ZooBot Austria 155: 213-232 pdf

Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510 pdf

Bosmans R, Van Keer K (2017) Een herziene soortenlijst van de Belgische spinnen (Araneae). Nieuwsbrief van de Belgische Arachnologische Vereniging 32: 39-69 pdf

Cera I (2018) The checklist of Latvian spiders (Arachnida: Araneae). Environmental and Experimental Biology 16: 139-152 pdf

Deltshev C, Vrenozi B, Blagoev G, Lazarov S (2011b) Spiders of Albania – faunistic and zoogeographical review (Arachnida: Araneae). Acta Zoologica Bulgarica 63: 125-144 pdf

Esyunin S L, Efimik V E (1996a) Catalogue of the spiders (Arachnida, Aranei) of the Urals. KMK Scientific Press, Moscow, 228 pp. pdf

Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp.

Geci D, Naumova M, Ibrahimi H, Grapci-Kotori L, Gashi A, Bilalli A, Musliu M (2023) Contribution to spider fauna (Arachnida: Araneae) from Bjeshkët e Nemuna mountains (Kosovo). Natura Croatica 32: 525-547 pdf

Helsdingen P J van (2025a) Catalogus van de Nederlandse spinnen. Versie 2025.1. 249 pp. pdf

Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204 pdf

Ivanov V V (2013) The checklist of Belarusian spiders (Arachnida, Araneae). Zoology and Ecology 4: 293-311 pdf

Koponen S, Fritzén N R, Pajunen T (2016) Checklist of spiders in Finland (Araneae). 6th version, December 2016, online at http://biolcoll.utu.fi/arach/checklist_of_spiders_in_Finland.htm pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Kreuels M, Staudt A, Christian S (2019) Die Spinnenfauna von Luxemburg – eine Zusammenstellung der Nachweise aus den Jahren 1906-2018 (Arachnida: Araneae). Bulletin de la Société des naturalistes luxembourgeois 121: 203-230 pdf

Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001] pdf

Lavery A (2019) A revised checklist of the spiders of Great Britain and Ireland. Arachnology 18: 196-212 pdf

Le Peru B (2007) Catalogue et répartition des araignées de France. Revue Arachnologique 16: 1-468 pdf

Lecigne S (2018b) Récits de chasses aranéologiques récentes dans plusieurs départements de France. Redécouverte de Philodromus buchari Kubcová, 2004 (Araneae: Philodromidae) et confirmation de la présence de Theridion harmsi Wunderlich, 2011 (Araneae: Theridiidae). Nieuwsbrief van de Belgische Arachnologische Vereniging 33: 59-99 pdf

Løvbrekke H (unpubl.)

Maurer R, Hänggi A (1990) Katalog der schweizerischen Spinnen. Documenta Faunistica Helvetiae 12 pdf

Naumova M, Lazarov S, Deltshev C (2019a) Faunistic diversity of the spiders in Montenegro (Arachnida: Araneae). Ecologica Montenegrina 22: 50-89 pdf

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Roberts M J (1987) The spiders of Great Britain and Ireland. Volume 2: Linyphiidae and checklist. Harley Books Colchester, England, 204 pp. pdf

Roberts M J (1995) Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp. pdf

Roberts M J (1998) Spinnengids. Tirion, Baarn, Netherlands, 397 pp. pdf

Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Scharff N, Gudik-Sørensen O (2006) Katalog over Danmarks edderkopper (Araneae). Catalogue of the spiders of Denmark (Araneae). Entomologiske Meddelelser 74: 3-71 pdf

Schikora H-B (2014) Die Webspinnen des Regenrnoorkomplexes Zehlaubruch, Kaliningrad-Region von Russland (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins zu Bremen 47: 427-446 pdf

Thaler K (1995) Beiträge zur Spinnenfauna von Nordtirol - 5. Linyphiidae 1: Linyphiinae (sensu Wiehle) (Arachnida: Araneida). Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck 82: 153-190 pdf

Vilbaste A (1987) Eesti Ämblikud (Aranei). Annoteeritud nimestik. Estonian spiders (Aranei). An annotated check list. Valgus, Tallinn. 113 pp. & 512 maps on 67 unpag. pages pdf

Vrenozi B (2012) A collection of spiders (Araneae) in Albanian coastal areas. Arachnologische Mitteilungen 44: 41-46 pdf

Weiss I, Petrișor A (1999) List of the spiders (Arachnida: Araneae) from Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 41: 79-107 pdf

Wiehle H (1956) Spinnentiere oder Arachnoidea, X. 28: Familie Linyphiidae-Baldachinspinnen. Die Tierwelt Deutschlands 44: 1-337 pdf

WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (13.2.2026) doi: 10.24436/2 pdf

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