Titanoecidae  Lehtinen, 1967 / Titanoeca  Thorell, 1870   17 species

Titanoeca quadriguttata (Hahn, 1833)

Description

Epigyne forms a narrow slit with parallel lateral margins. Prosoma reddish brown. Sternum black-brown. Chelicerae black-brown. Legs red-brown. Femur black. Opisthosoma almost black to brownish red, velvety-haired, dorsally with 2 pairs of large, white spots.

Body length male: 4.5-5 mm
Body length female: 4.5-5.5 mm
Additional information

Under stones on sunny, warm slopes

Distribution
  Presence
  No data
Global distribution (WSC 2026): Europe, Russia (Europe to South Siberia), Kazakhstan, China
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List

"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.

Albania   (Deltshev et al., 2011b; Vrenozi, 2012; Blick, 2018) ||| Austria   (Thaler, 1994b) ||| Azerbaijan   (Nuruyeva & Zamani, 2025) ||| Belgium   (Bosmans & Van Keer, 2017) ||| Bulgaria   (Blagoev et al., 2018) ||| Croatia   (Milosevic, 2002) ||| Czechia   (Růžička & Řezáč, 2022a) ||| France   (Le Peru, 2007) ||| Germany   (Blick et al., 2016) ||| Greece   (Bosmans & Chatzaki, 2005) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Isaia et al., 2025) ||| Liechtenstein   (Arnold, 2001) ||| Luxembourg   (Kreuels et al., 2019) ||| Moldova   (Vasiliev & Mikhailov, 2023) ||| Netherlands   (van Helsdingen, 2025a) ||| North Macedonia   (Komnenov, 2017) ||| Poland   (Rozwałka & Stanska, 2008) ||| Portugal   (Branco et al., 2019) ||| Romania   (Weiss & Petrișor, 1999) ||| Russia, Eastern   (Esyunin & Efimik, 1996a) ||| Russia, Southern   (Ponomarev, 2022) ||| Slovakia   (Gajdoš et al., 1999) ||| Slovenia   (Kostanjšek & Kuntner, 2015) ||| Spain   (Branco et al., 2019) ||| Switzerland   (Maurer & Hänggi, 1990) ||| Turkey (Europe)   (Demircan & Topçu, 2015; Demircan Aksan & Topçu, 2022c) ||| Ukraine   (Polchaninova & Prokopenko, 2019) |||
References

Arnold K (2001) Beitrag zur Spinnenfauna (Arachnida, Araneae) des Fürstentums Liechtenstein. Berichte der Botanisch-Zoologischen Gesellschaft Liechtenstein-Sargans-Werdenberg 28: 211-244 pdf

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Blick T (2018) A small collection of spiders (Arachnida: Araneae) from the River Vjosa, Albania – with an updated spider checklist of Albania. Acta ZooBot Austria 155: 213-232 pdf

Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510 pdf

Bosmans R, Chatzaki M (2005) A catalogue of the spiders of Greece – a critical review of all spider species cited from Greece with their localities. Nieuwsbrief van de Belgische arachnologische Vereniging 20 (2, suppl.): 1-124 pdf

Bosmans R, Van Keer K (2017) Een herziene soortenlijst van de Belgische spinnen (Araneae). Nieuwsbrief van de Belgische Arachnologische Vereniging 32: 39-69 pdf

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254 pdf

Deltshev C, Vrenozi B, Blagoev G, Lazarov S (2011b) Spiders of Albania – faunistic and zoogeographical review (Arachnida: Araneae). Acta Zoologica Bulgarica 63: 125-144 pdf

Demircan Aksan N, Topçu A (2022c) A preliminary checklist of the spider fauna of European Türkiye. Serket 19: 39-56 pdf

Demircan N, Topçu A (2015) A contribution to the spider fauna of the European part of Turkey (Araneae). Serket 14: 176-183 pdf

Esyunin S L, Efimik V E (1996a) Catalogue of the spiders (Arachnida, Aranei) of the Urals. KMK Scientific Press, Moscow, 228 pp. pdf

Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp.

Helsdingen P J van (2025a) Catalogus van de Nederlandse spinnen. Versie 2025.1. 249 pp. pdf

Hubert M (1966) Remarques sur quelques espèces d'araignées appartenant au genre Titanoeca Thorell 1870. Bulletin du Muséum National d'Histoire Naturelle de Paris 38: 238-246 pdf

Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204 pdf

Komnenov M (2017) New data on spider fauna (Araneae) of Shar Mountain, north-western Macedonia. Proceedings of the 5th Congress of the Ecologists of Macedonia, with international participation (Ohrid, 19th-22nd October 2016). Special issues of the Macedonian Ecological Society 13: 44-61 pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Kratochvíl J (1932b) Rod pavouku Titanoeca Thor. v Ceskoslovenske republice. Sborník Přírodovědecké Společnosti v Moravské Ostravě 7: 11-23 pdf

Kreuels M, Staudt A, Christian S (2019) Die Spinnenfauna von Luxemburg – eine Zusammenstellung der Nachweise aus den Jahren 1906-2018 (Arachnida: Araneae). Bulletin de la Société des naturalistes luxembourgeois 121: 203-230 pdf

Le Peru B (2007) Catalogue et répartition des araignées de France. Revue Arachnologique 16: 1-468 pdf

Loksa I (1969) Pokok I-Araneae I. Fauna Hungariae 97: 1-133 pdf

Maurer R, Hänggi A (1990) Katalog der schweizerischen Spinnen. Documenta Faunistica Helvetiae 12 pdf

Milosevic B (2002) Aranea – Popis vrsta – Checklist. Hrvatsko entomološko društvo – Inventar entomofaune Hrvatske [Aranea – List of species – Checklist. Croatian Entomological Society – Inventory of Croatian entomofauna]. pdf

Nuruyeva T, Zamani A (2025) An annotated checklist of the spiders (Arachnida: Araneae) of Azerbaijan. Zootaxa 5680: 1-168 pdf

Pesarini C (unpubl.)

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Roberts M J (1995) Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp. pdf

Rozwałka R, Stanska M (2008) Check-list of spiders (Araneae) of Poland. 1. December 2008. online at http://www.arachnologia.edu.pl/en/species/8-spiders-of-poland.html pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Thaler K (1994b) Beiträge zur Spinnenfauna von Nordtirol - 2. Orthognathe, cribellate und haplogyne Familien, Phocidae, Zodariidae, Mimetidae sowie Argiopiformia (ohne Linyphiidae s.l.) (Arachnida: Araneida). Mit Bemerkungen zur Spinnenfauna der Ostalpen. Veröffentlichungen des Museum Ferdinandeum in Innsbruck 73(1993): 69-119 pdf

Vasiliev A I, Mikhailov K G (2023) Annotated checklist of spiders (Araneae) of the Republic of Moldova. Invertebrate Zoology 20: 307-342 pdf

Vrenozi B (2012) A collection of spiders (Araneae) in Albanian coastal areas. Arachnologische Mitteilungen 44: 41-46 pdf

Weiss I (1979) Lessertinella carpatica sp. n. (Arachnida, Araneae, Erigonidae). Reichenbachia 17: 325-330 pdf

Weiss I, Petrișor A (1999) List of the spiders (Arachnida: Araneae) from Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 41: 79-107 pdf

Wiehle H (1953) Spinnentiere oder Arachnoidea (Araneae), IX. Orthognatha-Cribellata-Haplogynae, Entelegynae (Pholcidae, Zodariidae, Oxyopidae, Mimetidae, Nesticidae). Die Tierwelt Deutschlands 42: 1-150 pdf

WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (13.2.2026) doi: 10.24436/2 pdf

Updates
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