Theridiidae  Sundevall, 1833 / Crustulina  Menge, 1868   6 species

Crustulina guttata  (Wider, 1834)

Description

Cymbium distally broad. Epigyne with lateral grooves. Prosoma dark brown, sometimes bright red-brown. Sternum dark brown. Chelicerae dark brown. Legs dark brown, vaguely annulated, short and robust. Opisthosoma almost circular, brown, dorsally with 3 longitudinal rows of white spots (rarely missing).

Body length male: 2-2.5 mm
Body length female: 2.0-2.5 mm
Additional information

On the ground of open forest areas

Distribution
  Presence
  No data
Global distribution (WSC 2021): Canary Is., Europe, Caucasus, Russia (Europe to South Siberia), Kazakhstan, Iran, Central Asia, China, Korea, Japan
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List
Albania   (Deltshev et al., 2011b) ||| Austria   (no references) ||| Azerbaijan   (Otto, 2020) ||| Belarus   (Ivanov, 2013) ||| Belgium   (no references) ||| Bulgaria   (Blagoev et al., 2018) ||| Croatia   (no references) ||| Czechia   (no references) ||| Denmark   (no references) ||| Estonia   (no references) ||| Finland   (Koponen et al., 2016) ||| France   (no references) ||| France / Corsica   (no references) ||| Georgia   (Otto, 2020) ||| Germany   (no references) ||| Greece   (no references) ||| Hungary   (Samu & Szinetár, 1999) ||| Ireland   (Nolan, 2014) ||| Italy   (Pantini & Isaia, 2019) ||| Italy / Sicily   (Dentici & Galasso, 2021) ||| Kosovo   (Geci & Naumova, 2021b) ||| Latvia   (Cera, 2018) ||| Liechtenstein   (no references) ||| Lithuania   (no references) ||| Luxembourg   (Kreuels et al., 2019) ||| Moldova   (no references) ||| Montenegro   (no references) ||| Netherlands   (no references) ||| North Macedonia   (no references) ||| Norway   (Aakra & Hauge, 2003) ||| Poland   (no references) ||| Portugal   (Branco et al., 2019) ||| Romania   (no references) ||| Russia, Central   (no references) ||| Russia, Eastern   (no references) ||| Russia, Kaliningrad Region   (no references) ||| Russia, Northern   (no references) ||| Russia, Southern   (Otto, 2020) ||| Russia, Western   (no references) ||| Serbia   (no references) ||| Slovakia   (no references) ||| Slovenia   (Kostanjšek & Kuntner, 2015) ||| Spain   (Branco et al., 2019) ||| Sweden   (Kronestedt, 2001) ||| Switzerland   (no references) ||| Turkey (Europe)   (no references) ||| Ukraine   (Polchaninova & Prokopenko, 2019) ||| United Kingdom   (no references) |||
References

Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129 pdf

Azheganova N S (1968) Kratkii opredelitel' paukov (Aranei) lesnoi i lesostepnoi zony SSSR. Akademia Nauk SSSR, 149pp. pdf

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254 pdf

Cera I (2018) The checklist of Latvian spiders (Arachnida: Araneae). Environmental and Experimental Biology 16: 139-152 pdf

Deltshev C, Vrenozi B, Blagoev G, Lazarov S (2011b) Spiders of Albania – faunistic and zoogeographical review (Arachnida: Araneae). Acta Zoologica Bulgarica 63: 125-144 pdf

Dentici A, Galasso P (2021) Checklist of spiders and Opiliones (Arachnida Araneae, Opiliones) of Sicilian southest swamp lakes "Pantano Cuba", "Pantano Longarini" and “Pantano Bruno” (Italy). Fifth contribution to knowledge of Sicilian spider fauna Biodiversity Journal 21: 695-703 pdf

Geci D, Naumova M (2021b) A preliminary checklist of the spiders of Kosovo (Arachnida: Araneae). Ecologia Balkanica, Special Edition 4: 11-28 pdf

Helsdingen P J van (2013) A quick scan of the spider fauna of the European part of Turkey. Nieuwsbrief SPINED 33: 29-38 pdf

Ivanov V V (2013) The checklist of Belarusian spiders (Arachnida, Araneae). Zoology and Ecology 4: 293-311 pdf

Knoflach B (1994) Zur Genitalmorphologie und Biologie der Crustulina-Arten Europas (Arachnida: Araneae, Theridiidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 67: 327-346 pdf

Koponen S, Fritzén N R, Pajunen T (2016) Checklist of spiders in Finland (Araneae). 6th version, December 2016, online at http://biolcoll.utu.fi/arach/checklist_of_spiders_in_Finland.htm pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Kreuels M, Staudt A, Christian S (2019) Die Spinnenfauna von Luxemburg – eine Zusammenstellung der Nachweise aus den Jahren 1906-2018 (Arachnida: Araneae). Bulletin de la Société des naturalistes luxembourgeois 121: 203-230 pdf

Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001] pdf

Locket G H, Millidge A F (1953) British spiders. Vol. II. Ray Society, London, 449 pp. pdf

Nolan M (2014) First Irish records of Crustulina guttata (Wider) with notes on Irish occurrences of Crustulina sticta (O.P.-Cambridge) (Araneae: Theridiidae). Bulletin of the Irish Biogeographical Society 38: 285-290 pdf

Otto S (2020) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 03.2020. Internet: caucasus-spiders.info.

Pantini P, Isaia M (2019) Araneae.it: the online catalog of Italian spiders, with addenda on other arachnid orders occurring in Italy (Arachnida: Araneae, Opiliones, Palpigradi, Pseudoscorpionida, Scorpiones, Solifugae). Fragmenta Entomologica 51: 127-152 pdf

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Roberts M J (1995) Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp. pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Schröder M, Chatzaki M, Buchholz S (2011) The spider fauna of the Aladjagiola wetland complex (Nestos Delta, north-east Greece): a reflection of a unique zoogeographical transition zone in Europe. Biological Journal of the Linnean Society 102: 217-233 pdf

Wiehle H (1937) Spinnentiere oder Arachnoidea, VIII. 26: Familie Theridiidae oder Haubennetzspinnen (Kugelspinnen). Die Tierwelt Deutschlands 33: 119-222 pdf

WSC (2021) World Spider Catalog. Version 22.5. Natural History Museum Bern, online at http://wsc.nmbe.ch (22.09.2021) doi: 10.24436/2 pdf

Yoshida H (2001) The spider genera Robertus, Enoplognatha, Steatoda and Crustulina (Araneae: Theridiidae) from Japan. Acta Arachnologica 50: 31-48 pdf

Zamani A, Dimitrov D, Weiss I, Alimohammadi S, Rafiei-Jahed R, Esyunin S L, Moradmand M, Chatzaki M, Marusik Y M (2020c) New data on the spider fauna of Iran (Arachnida: Araneae), part VII. Arachnology 18: 569-591 pdf

Updates
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