Gnaphosidae  Banks, 1892 / Civizelotes  Senglet, 2012   13 species

Civizelotes gracilis (Canestrini, 1868)

Description

Epigyne with cuticular bar straightly converging posteriorly. Prosoma black-brown to black. Legs brighter. Opisthosoma black-brown to black, with weakly pronounced scuticula.

Body length male: 2.0 mm
Body length female: 2.2-2.5 mm
Additional information

On dry, south-exposed slopes and rock steppes

Distribution
  Presence
  No data
Global distribution (WSC 2026): Central and South-Eastern Europe, Caucasus (Russia, Georgia), Turkey
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List
Albania   (Naumova et al., 2016b) ||| Austria   (Thaler & Knoflach, 2004b) ||| Bulgaria   (Blagoev et al., 2018) ||| Croatia   (Milosevic, 2002) ||| Czechia   (Růžička & Řezáč, 2022a) ||| Georgia   (Otto, 2022) ||| Greece   (Schröder et al., 2011) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Isaia et al., 2025) ||| Moldova   (Vasiliev & Mikhailov, 2023) ||| North Macedonia   (Komnenov, 2014) ||| Romania   (Weiss & Petrișor, 1999) ||| Russia, Southern   (Otto, 2022; Ponomarev, 2022) ||| Serbia   (Grbić & Savić, 2010; Dudić et al., 2013; Grbić et al., 2015) ||| Slovakia   (Gajdoš et al., 1999) ||| Slovenia   (Kostanjšek & Kuntner, 2015) ||| Turkey (Asia)   (Danışman et al., 2025a) ||| Ukraine   (Polchaninova & Prokopenko, 2019) |||
References

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Danışman T, Kunt K B, Özkütük R S, Coşar İ (2025a) The checklist of the spiders of Turkey. Version 2025 [last updated 01 December 2025], online at http://www.spidersofturkey.info pdf

Deltshev C (2003) A critical review of the spider species (Araneae) described by P. Drensky in the period 1915-1942 from the Balkans. Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck 90: 135-150 pdf

Drensky P (1929) Paiatzi (Aranea) ot tzentralna i iougo-zapadna Makedonia. Spisanié na Beulgarskata Akademia na Naoukite 39: 1-76 pdf

Dudić B D, Tomić V T, Sivčev I, Büchs W, Sivčev L, Graora D, Gotlin-Čuljak T (2013) New data on spider fauna from northern Serbia. Archives of Biological Sciences 65: 1669-1673 pdf

Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp.

Grbić G, Hänggi A, Savić D (2015) New faunistic records of spiders (Arachnida, Areneae [sic]) from the Fruška Gora mountain, northern Serbia. Acta Zoologica Bulgarica 67: 479-486 pdf

Grbić G, Savić D (2010) Contribution to the knowledge of the spider fauna (Arachnida, Araneae) on the Fruška Gora Mt. Acta Entomologica Serbica 15: 243-260 pdf

Grimm U (1985) Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF) 26: 1-318 pdf

Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204 pdf

Komnenov M (2014) Spider fauna of the Osogovo Mt. Range, Northeastern Macedonia. Fauna Balkana 2: 1-267 pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Miller F (1967) Studien über die Kopulationsorgane der Spinnengattung Zelotes, Micaria, Robertus und Dipoena nebst Beschreibung einiger neuen oder unvollkommen bekannten Spinnenarten. Přírodovědné práce ústavů Československé Akademie Věd v Brně (N.S.) 1: 251-298 pdf

Milosevic B (2002) Aranea – Popis vrsta – Checklist. Hrvatsko entomološko društvo – Inventar entomofaune Hrvatske [Aranea – List of species – Checklist. Croatian Entomological Society – Inventory of Croatian entomofauna]. pdf

Naumova M, Hristovski S, Hristov G H (2016b) Spiders (Arachnida: Araneae) from Prespa National Park, Albania. Acta Zoologica Bulgarica 68: 503-511 pdf

Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Schröder M, Chatzaki M, Buchholz S (2011) The spider fauna of the Aladjagiola wetland complex (Nestos Delta, north-east Greece): a reflection of a unique zoogeographical transition zone in Europe. Biological Journal of the Linnean Society 102: 217-233 pdf

Thaler K, Knoflach B (2004b) Zur Faunistik der Spinnen (Araneae) von Österreich: Gnaphosidae, Thomisidae (Dionycha pro parte). Linzer Biologische Beiträge 36: 417-484 pdf

Vasiliev A I, Mikhailov K G (2023) Annotated checklist of spiders (Araneae) of the Republic of Moldova. Invertebrate Zoology 20: 307-342 pdf

Weiss I, Petrișor A (1999) List of the spiders (Arachnida: Araneae) from Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 41: 79-107 pdf

Weiss I, Sarbu S (1977) Zur Kenntnis der Spinnen und Weberknechte des Botanischen Gartens Iasi. Studii şi Comunicări Muzeul Brukenthal, Ştiin ţe Naturale 21: 225-243 pdf

WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (13.2.2026) doi: 10.24436/2 pdf

Updates
03-02-2026 Distribution update Detail
03-02-2026 Distribution insert Detail
23-07-2025 Distribution update Detail
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30-09-2015 Distribution update Detail