Heser nilicola (O. Pickard-Cambridge, 1874)
Description
Male
Embolus enormously elongated.
Body length male: 2.8-3.7 mmFemale
Massive epigynal ducts.
Body length female: 3.1-3.8 mmAdditional information
also in citrus groves
Distribution
Figures
Pedipalp, ventral view
(Platnick & Shadab 1983)
Pedipalp, retrolateral view
(Platnick & Shadab 1983)
Pedipalp, prolateral-ventral view
(Chatzaki et al. 2003)
Pedipalp, retrolateral view
(Chatzaki et al. 2003)
Pedipalp, prolateral view
(Senglet 2012)
Pedipalp, ventral view
(Senglet 2012)
Pedipalp, retrolateral view
(Senglet 2012)
Pedipalp, retrolateral view
(Cornic et al. 2004)
Pedipalp, retrolateral view
(Levy 1998c)
Pedipalp, ventral view
(Levy 1998c)
Pedipalp, retrolateral view
(FitzPatrick 2007)
Pedipalp, ventral view
(FitzPatrick 2007)
Bulbus, dorsal view
(Chatzaki et al. 2003)
Epigyne
(Platnick & Shadab 1983)
Epigyne, ventral view
(Senglet 2012)
Epigyne, ventral view
(Senglet 2012)
Epigyne
(Danışman et al. 2024b)
Epigyne
(Cornic et al. 2004)
Epigyne
(Levy 1998c)
Epigyne
(FitzPatrick 2007)
Vulva, dorsal view
(Platnick & Shadab 1983)
Vulva, ventral view
(Chatzaki et al. 2003)
Vulva
(Chatzaki et al. 2003)
Vulva, dorsal view
(Senglet 2012)
Vulva, dorsal view
(Danışman et al. 2024b)
Vulva, dorsal view
(Levy 1998c)
Vulva, dorsal view
(FitzPatrick 2007)
Distribution List
"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.
References
Barrientos J A, Febrer B (2017) Arañas (Arachnida, Araneae) de Menorca (Islas Baleares, España). Nuevos datos. Revista Ibérica de Aracnología 31: 8-24 
Bosmans R, Van Keer J, Russell-Smith A, Hadjiconstantis M, Komnenov M, Bosselaers J, Huber S, McCowan D, Snazell R, Decae A, Zoumides C, Kielhorn K-H, Oger P (2019c) Spiders of Cyprus (Araneae). A catalogue of all currently known species from Cyprus. Newsletter of the Belgian Arachnological Society 34, Suppl. 1: 1-173
Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254
Chatzaki M, Thaler K, Mylonas M (2003) Ground spiders (Gnaphosidae; Araneae) from Crete and adjacent areas of Greece. Taxonomy and distribution. III. Zelotes and allied genera. Revue Suisse de Zoologie 110: 45-89
Cornic J-F, Ledoux J-C, Villepoux O (2004) De araneis Galliae I.2, Zelotes nilicola O. P.-Cambridge. Revue Arachnologique 17
Danışman T, Ülker A, Coşar İ (2024b) Genus Heser Tuneva, 2004 (Araneae: Gnaphosidae), a new record for the spider fauna of Türkiye. Munis Entomology and Zoology 19: 1029-1031
El-Hennawy H K (2017a) A list of Egyptian spiders (revised in 2017). Serket 15: 167-183
FitzPatrick M J (2007) A taxonomic revision of the Afrotropical species of Zelotes (Arachnida: Araneae: Gnaphosidae). Bulletin of the British Arachnological Society 14: 97-172
Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204
Kmira G, Nasri-Ammar, Petillon J, Bosmans R (2025) Catalog of the spiders of Tunisia (Arachnida: Araneae). Zootaxa 5641: 1-95
Le Peru B (2007) Catalogue et répartition des araignées de France. Revue Arachnologique 16: 1-468
Levy G (1998c) The ground-spider genera Setaphis, Trachyzelotes, Zelotes, and Drassyllus (Araneae: Gnaphosidae) in Israel. Israel Journal of Zoology 44: 93-158
Mestre L, Piñol J, Barrientos J A, Cama A, Espadaler X (2012) Effects of ant competition and bird predation on the spider assemblage of a citrus grove. Basic and Applied Ecology 13: 355-362
Pfliegler W A, Schönhofer A, Niedbała W, Vella P, Sciberras A, Vella A (2017) New records of mites (Acari) and harvestmen (Opiliones) from Malta with a preliminary checklist of Maltese Arachnida. Soil Organisms 89: 85-110
Platnick N I, Shadab M U (1983) A revision of the American spiders of the genus Zelotes (Araneae, Gnaphosidae). Bulletin of the American Museum of Natural History 174: 97-192
Senglet A (2012) Civizelotes new genus, and other new or little known Zelotinae (Araneae, Gnaphosidae). Revue Suisse de Zoologie 119: 501-528
WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (13.2.2026) doi: 10.24436/2