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Gnaphosidae  Banks, 1892 / Leptopilos  Levy, 2009   6 species

Leptopilos memorialis (Spassky, 1940)

WSC LSID urn:lsid:nmbe.ch:spidersp:027463

Key to genera | Key to species

Description
Male

Prosoma and sternum corn-coloured, sternum with dark brown margins. Opisthosoma pale grey. Legs yellow.

Prosoma length: 1.1 mm, prosoma width: 0.9 mm.

Body length male: 2.58-2.65 mm
Female

Colouration as in male.

Prosoma length: 1.2 mm, prosoma width: 1.0 mm.

Body length female: 2.83-3.55 mm
Distribution
  Presence
  No data
Global distribution (WSC 2026): Bulgaria, Greece, Ukraine, Russia (Europe to Central Asia), Azerbaijan, Kazakhstan, Pakistan, Mongolia
Figures
species

male Pedipalp, ventral view
(Kovblyuk & Nadolny 2010)

species

male Pedipalp, ventral view
(Kovblyuk & Nadolny 2010)

species

male Pedipalp, retrolateral-ventral view
(Kovblyuk & Nadolny 2010)

species

male Pedipalp, prolateral view
(Kovblyuk & Nadolny 2010)

species

male Pedipalp, prolateral view
(Naumova et al. 2021)

species

male Pedipalp, retrolateral view
(Naumova et al. 2021)

species

male Pedipalp, ventral view
(Naumova et al. 2021)

species

male Eyes, frontal view
(Kovblyuk & Nadolny 2010)

species

female Epigyne
(Kovblyuk & Nadolny 2010)

species

female Epigyne
(van Helsdingen et al. 2018)

species

female Epigyne
(Naumova et al. 2021)

species

female Vulva, ventral view
(Kovblyuk & Nadolny 2010)

species

female Vulva, dorsal view
(Kovblyuk & Nadolny 2010)

species

female Vulva, dorsal view
(Naumova et al. 2021)

species

female Eyes, frontal view
(Kovblyuk & Nadolny 2010)

species

female Ocular area, lateral view
(Kovblyuk & Nadolny 2010)

Distribution List

"No references" does not mean that the species does not occur in this country, but that we have not yet inserted the reference for it. We are working on it.

Azerbaijan   (Nuruyeva & Zamani, 2025) ||| Bulgaria   (Naumova et al., 2021) ||| Greece   (van Helsdingen et al., 2018) ||| Russia, Eastern   (Tuneva & Esyunin, 2002) ||| Russia, Southern   (Otto, 2022; Ponomarev, 2022) ||| Ukraine   (Polchaninova & Prokopenko, 2019) |||
References

Helsdingen P J van, IJland S, Komnenov M (2018) Inventory of the spiders (Araneae) of the border region of northern Greece with Albania and FYR Macedonia. Nieuwsbrief SPINED 37: 5-23 pdf

Kovblyuk M M, Nadolny A A (2010) Cryptodrassus hungaricus and Leptodrassex memorialis from Crimea (Aranei: Gnaphosidae). Arthropoda Selecta 19: 189-197 pdf

Naumova M, Blagoev G, Deltshev D (2021) Fifty spider species new to the Bulgarian fauna, with a review of some dubious species (Arachnida: Araneae). Zootaxa 4984: 228-257 pdf

Nuruyeva T, Zamani A (2025) An annotated checklist of the spiders (Arachnida: Araneae) of Azerbaijan. Zootaxa 5680: 1-168 pdf

Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Tuneva T K, Esyunin S L (2002) A review of the family Gnaphosidae in the fauna of the Urals (Aranei), 2. New and rare genera. Arthropoda Selecta 10: 217-224 pdf

WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (13.2.2026) doi: 10.24436/2 pdf

Updates
19-08-2025 Distribution insert Detail
25-04-2025 Distribution update Detail
12-07-2021 Distribution update Detail
12-06-2021 Image insert
10-06-2021 Distribution insert Detail
17-03-2018 Image insert
17-03-2018 Distribution update Detail
28-09-2016 Datasheet update Detail
01-12-2013 Distribution update Detail