Gnaphosidae  Banks, 1892 / Drassodes  Westring, 1851   44 species

Drassodes villosus (Thorell, 1856)

Description

Male palp with distal apophysis, hook-shaped. Tibial apophysis with serrated margin. Embolus small. Epigyne with trapezoid central area, wider than long. Prosoma brown, eye region darker. Chelicerae darker than prosoma. Opisthosoma brown.

Body length male: 10.1-11.5 mm
Body length female: 10.5-15.2 mm
Additional information

In dry pine forests, under stones and bark during the day

Distribution
  Presence
  No data
Global distribution (WSC 2026): Europe, Turkey, Russia (Europe to Far East), Iran, Central Asia
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List
Austria   (Thaler & Knoflach, 2004b) ||| Bulgaria  Removed (Blagoev et al., 2018; Naumova et al., 2021) ||| Estonia   (Mikhailov, 2013) ||| Finland   (Koponen et al., 2016) ||| France   (Le Peru, 2007) ||| Georgia   (Otto, 2022) ||| Germany   (Blick et al., 2016) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Isaia et al., 2025) ||| Kosovo   (Geci et al., 2023) ||| Liechtenstein   (Arnold, 2001) ||| Lithuania   (Biteniekytė & Rėlys, 2012) ||| North Macedonia   (Deltshev et al., 2013) ||| Norway   (Aakra & Hauge, 2003) ||| Romania   (Weiss & Petrișor, 1999) ||| Russia, Central   (Ponomarev, 2022) ||| Russia, Eastern   (Esyunin & Efimik, 1996a) ||| Russia, Northern   (no references) ||| Russia, Southern   (Otto, 2022; Ponomarev, 2022; Ponomarev & Khnykin, 2013) ||| Russia, Western   (no references) ||| Serbia   (Naumova et al., 2021) ||| Slovakia   (Prídavka, 2002) ||| Spain / Balearic Islands   (Barrientos & Febrer, 2017) ||| Sweden   (Kronestedt, 2001) ||| Switzerland   (Maurer & Hänggi, 1990) ||| Turkey (Asia)   (Danışman et al., 2025a) ||| Ukraine   (Polchaninova & Prokopenko, 2019) |||
References

Aakra K, Hauge E (2003) Checklist of Norwegian spiders (Arachnida: Araneae), including Svalbard and Jan Mayen. Norwegian Journal of Entomology 50: 109-129 pdf

Almquist S (2007) Swedish Araneae, part 2 – families Dictynidae to Salticidae. Insect Systematics & Evolution, Supplement 63(2006): 285-601 pdf

Arnold K (2001) Beitrag zur Spinnenfauna (Arachnida, Araneae) des Fürstentums Liechtenstein. Berichte der Botanisch-Zoologischen Gesellschaft Liechtenstein-Sargans-Werdenberg 28: 211-244 pdf

Azheganova N S (1968) Kratkii opredelitel' paukov (Aranei) lesnoi i lesostepnoi zony SSSR. Akademia Nauk SSSR, 149pp. pdf

Barrientos J A, Febrer B (2017) Arañas (Arachnida, Araneae) de Menorca (Islas Baleares, España). Nuevos datos. Revista Ibérica de Aracnología 31: 8-24 pdf

Biteniekytė M, Rėlys V (2012) The checklist of Lithuanian spiders (Arachnida: Araneae). Biologija 57: 148-158 pdf

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510 pdf

Danışman T, Kunt K B, Özkütük R S, Coşar İ (2025a) The checklist of the spiders of Turkey. Version 2025 [last updated 01 December 2025], online at http://www.spidersofturkey.info pdf

Deltshev C, Komnenov M, Blagoev G, Georgiev T, Lazarov S, Stojkoska E, Naumova M (2013) Faunistic diversity of spiders (Araneae) in Galichitsa Mountain (FYR Macedonia). Biodiversity Data Journal 1(e977): 1-70 pdf

Esyunin S L, Efimik V E (1996a) Catalogue of the spiders (Arachnida, Aranei) of the Urals. KMK Scientific Press, Moscow, 228 pp. pdf

Geci D, Naumova M, Ibrahimi H, Grapci-Kotori L, Gashi A, Bilalli A, Musliu M (2023) Contribution to spider fauna (Arachnida: Araneae) from Bjeshkët e Nemuna mountains (Kosovo). Natura Croatica 32: 525-547 pdf

Grimm U (1985) Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF) 26: 1-318 pdf

Heimer S, Nentwig W (1991) Spinnen Mitteleuropas. Paul Parey Berlin pdf

Hervé C, Rollard C (2009) Drassodes species from the Parc national du Mercantour (French Alps), with the description of a new species (Araneae: Gnaphosidae). Contributions to Natural History 12: 627-642 pdf

Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204 pdf

Koponen S, Fritzén N R, Pajunen T (2016) Checklist of spiders in Finland (Araneae). 6th version, December 2016, online at http://biolcoll.utu.fi/arach/checklist_of_spiders_in_Finland.htm pdf

Kronestedt T (2001) Checklist of Spiders (Araneae) in Sweden [Preliminary version February 2001] pdf

Le Peru B (2007) Catalogue et répartition des araignées de France. Revue Arachnologique 16: 1-468 pdf

Løvbrekke H (unpubl.)

Maurer R, Hänggi A (1990) Katalog der schweizerischen Spinnen. Documenta Faunistica Helvetiae 12 pdf

Mikhailov K G (2013) The spiders (Arachnida: Aranei) of Russia and adjacent countries: a non-annotated checklist. Arthropoda Selecta, Supplement 3: 1-262 pdf

Naumova M, Blagoev G, Deltshev D (2021) Fifty spider species new to the Bulgarian fauna, with a review of some dubious species (Arachnida: Araneae). Zootaxa 4984: 228-257 pdf

Otto S (2022) Caucasian spiders. A faunistic database on the spiders of the Caucasus Ecoregion. Database version 02.2022. Internet: caucasus-spiders.info.

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Ponomarev A V, Khnykin A S (2013) Spiders (Aranei) of Volgograd sity and its environs. The South of Russia: Ecology, Development 2013(4): 109-136 pdf

Prídavka R (2002) Príspevok k poznaniu pavúkov (Araneae) Borskej nížiny. [Contribution to the knowledge of spiders (Araneae) of Borská Nižina lowland]. Sborník Přirodovědného klubu v Uherském Hradišti 7: 91-104

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Thaler K, Knoflach B (2004b) Zur Faunistik der Spinnen (Araneae) von Österreich: Gnaphosidae, Thomisidae (Dionycha pro parte). Linzer Biologische Beiträge 36: 417-484 pdf

Tullgren A (1946) Svensk spindelfauna. 3. Egentiliga spinddlar. Araneae Fam 5-7. Clubionidae, Zoridae och Gnaphosidae. Stockholm (Entomol Fören.): 141 pp. pdf

Weiss I, Petrișor A (1999) List of the spiders (Arachnida: Araneae) from Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 41: 79-107 pdf

WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (20.5.2026) doi: 10.24436/2 pdf

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