Gnaphosidae  Banks, 1892 / Drassyllus  Chamberlin, 1922   19 species

Drassyllus villicus (Thorell, 1875)

Description

Tibial apophysis distally forked. Epigyne with interrupted anterior margin. Prosoma dark red-brown. Legs brighter than prosoma. Femur I with bright spot. Opisthosoma dark red-brown with dark brown scuticula.

Body length male: 4.5-4.9 mm
Body length female: 5.1-5.8 mm
Additional information

In warm localities (open shrubbery and forests, steppes, dry meadows)

Distribution
  Presence
  No data
Global distribution (WSC 2026): Europe, Turkey, Azerbaijan
Phenology
JanFebMarAprMayJunJulAugSepOctNovDec
 male  female
Figures
Distribution List
Albania   (Vrenozi & Jäger, 2012) ||| Austria   (Thaler & Knoflach, 2004b) ||| Azerbaijan   (Khasayeva & Huseynov, 2017) ||| Belgium   (Bosmans & Van Keer, 2017) ||| Bosnia and Herzegovina   (Kuralt et al., 2025) ||| Bulgaria   (Blagoev et al., 2018) ||| Croatia   (Milosevic, 2002) ||| Czechia   (Růžička & Řezáč, 2022a) ||| France   (Le Peru, 2007) ||| Germany   (Blick et al., 2016) ||| Greece   (Van Keer et al., 2010) ||| Hungary   (Samu & Szinetár, 1999) ||| Italy   (Isaia et al., 2025) ||| Kosovo   (Geci & Naumova, 2021b) ||| Moldova   (Vasiliev & Mikhailov, 2023) ||| North Macedonia   (Komnenov, 2014) ||| Portugal   (Branco et al., 2019) ||| Romania   (Weiss & Petrișor, 1999) ||| Russia, Central   (Ponomarev, 2022) ||| Serbia   (Grbić et al., 2015) ||| Slovakia   (Gajdoš et al., 1999) ||| Slovenia   (Kostanjšek & Kuntner, 2015; Gregorič & Kuntner, 2009) ||| Spain   (Branco et al., 2019) ||| Switzerland   (Maurer & Hänggi, 1990) ||| Turkey (Asia)   (Danışman et al., 2025a; Seyyar et al., 2008a) ||| Ukraine   (Polchaninova & Prokopenko, 2019) |||
References

Blagoev G, Deltshev C, Lazarov S, Naumova M (2018) The spiders (Araneae) of Bulgaria. Version: August 2018. National Museum of Natural History, Bulgarian Academy of Sciences. Online at http://www.nmnhs.com/spiders-bulgaria/ (accessed on 10.9.2018) pdf

Blick T, Finch O-D, Harms K H, Kiechle J, Kielhorn K-H , Kreuels M, Malten A, Martin D, Muster C, Nährig D, Platen R, Rödel I, Scheidler M, Staudt A, Stumpf H, Tolke D (2016) Rote Liste und Gesamtartenliste der Spinnen (Arachnida: Araneae) Deutschlands. 3. Fassung, Stand April 2008, einzelne Änderungen und Nachträge bis August 2015. Naturschutz und Biologische Vielfalt 70/4: 383-510 pdf

Bosmans R, Van Keer K (2017) Een herziene soortenlijst van de Belgische spinnen (Araneae). Nieuwsbrief van de Belgische Arachnologische Vereniging 32: 39-69 pdf

Branco V V, Morano E, Cardoso P (2019) An update to the Iberian spider checklist (Araneae). Zootaxa 4614: 201-254 pdf

Danışman T, Kunt K B, Özkütük R S, Coşar İ (2025a) The checklist of the spiders of Turkey. Version 2025 [last updated 01 December 2025], online at http://www.spidersofturkey.info pdf

Gajdoš P, Svatoň J, Sloboda J (1999) Katalóg pavúkov Slovenska. Catalogue of Slovakian spiders. Ústav kr. ekol. SAV, Bratislava, 337 pp.

Geci D, Naumova M (2021b) A preliminary checklist of the spiders of Kosovo (Arachnida: Araneae). Ecologia Balkanica, Special Edition 4: 11-28 pdf

Grbić G, Hänggi A, Savić D (2015) New faunistic records of spiders (Arachnida, Areneae [sic]) from the Fruška Gora mountain, northern Serbia. Acta Zoologica Bulgarica 67: 479-486 pdf

Gregorič M, Kuntner M (2009) Epigean spider diversity in the classical karst. Hacquetia 8: 67-78 pdf

Grimm U (1985) Die Gnaphosidae Mitteleuropas (Arachnida, Araneae). Abhandlungen des Naturwissenschaftlichen Vereins in Hamburg (NF) 26: 1-318 pdf

Isaia M, Tolve M, Pantini P (2025) Araneae.it: 2.0: the new updated version of the digital catalog of the Italian spiders and other arachnids (Arachnida: Araneae, Opiliones, Scorpiones, Pseudoscorpiones, Palpigradi, Solifugae). Fragmenta Entomologica 57: 197-204 pdf

Khasayeva S I, Huseynov E F (2017) To the study of gnaphosid spiders (Aranei: Gnaphosidae) of Western Azerbaijan with suggestion of two new synonyms. Arthropoda Selecta 26: 125-132 pdf

Komnenov M (2014) Spider fauna of the Osogovo Mt. Range, Northeastern Macedonia. Fauna Balkana 2: 1-267 pdf

Kostanjšek R, Kuntner M (2015) Araneae Sloveniae: a national spider species checklist. ZooKeys 474: 1-91 pdf

Kovblyuk M M (2003b) The spider genus Drassyllus Chamberlin, 1922 in the Crimean fauna, with description of a new species (Aranei: Gnaphosidae). Arthropoda Selecta 12: 23-28 pdf

Kuralt Ž, Pajek Arambašič N, Kostanjšek R (2025) Contribution to the knowledge of spiders (Araneae) of the upper course of the Neretva River in Bosnia and Herzegovina. Natura Sloveniae 27: 114-122 pdf

Le Peru B (2007) Catalogue et répartition des araignées de France. Revue Arachnologique 16: 1-468 pdf

Maurer R, Hänggi A (1990) Katalog der schweizerischen Spinnen. Documenta Faunistica Helvetiae 12 pdf

Miller F (1967) Studien über die Kopulationsorgane der Spinnengattung Zelotes, Micaria, Robertus und Dipoena nebst Beschreibung einiger neuen oder unvollkommen bekannten Spinnenarten. Přírodovědné práce ústavů Československé Akademie Věd v Brně (N.S.) 1: 251-298 pdf

Milosevic B (2002) Aranea – Popis vrsta – Checklist. Hrvatsko entomološko društvo – Inventar entomofaune Hrvatske [Aranea – List of species – Checklist. Croatian Entomological Society – Inventory of Croatian entomofauna]. pdf

Polchaninova N, Prokopenko E (2019) An updated checklist of spiders (Arachnida: Araneae) of Left-Bank Ukraine. Arachnologische Mitteilungen 57: 60-64 & App pdf

Ponomarev A V (2022) Spiders (Arachnida: Araneae) of the southeast of the Russian Plain: catalogue, the fauna specific features. SSC RAS Publishers, Rostov-on-Don, 640 pp. pdf

Roberts M J (1995) Collins Field Guide: Spiders of Britain & Northern Europe. HarperCollins, London, 383 pp. pdf

Růžička V, Řezáč M (2022a) Seznam pavouků České republiky. List of spiders of the Czech Republic. Online at https://www.arachnology.cz/seznam-pavouku-cr-26.html and https://www.arachnology.cz/en/seznam-pavouku-cr-26.html pdf

Samu F, Szinetár C (1999) Bibliographic check list of the Hungarian spider fauna. Bulletin of the British Arachnological Society 11: 161-184 pdf

Seyyar O, Ayyildiz N, Topçu A (2008a) Updated checklist of ground spiders (Araneae: Gnaphosidae) of Turkey, with zoogeographical and faunistic remarks. Entomological News 119: 509-520 pdf

Thaler K, Knoflach B (2004b) Zur Faunistik der Spinnen (Araneae) von Österreich: Gnaphosidae, Thomisidae (Dionycha pro parte). Linzer Biologische Beiträge 36: 417-484 pdf

Van Keer J, Van Keer K, De Koninck H, Ramel G (2010) The arachnofauna (Araneae) of wetland Kerkini (Macedonia-Northern Central Greece). Acta Zoologica Bulgarica 62: 141-160 pdf

Vasiliev A I, Mikhailov K G (2023) Annotated checklist of spiders (Araneae) of the Republic of Moldova. Invertebrate Zoology 20: 307-342 pdf

Vrenozi B, Jäger P (2012) A faunistic study on ground-dwelling spiders (Araneae) in the Tirana district, Albania. Arachnologische Mitteilungen 44: 81-87 pdf

Weiss I, Petrișor A (1999) List of the spiders (Arachnida: Araneae) from Romania. Travaux du Muséum National d’Histoire Naturelle “Grigore Antipa” 41: 79-107 pdf

WSC (2026) World Spider Catalog. Version 27. Natural History Museum Bern, online at http://wsc.nmbe.ch (13.2.2026) doi: 10.24436/2 pdf

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